Stuttgarter Beiträge zur Naturkunde Serie A (Biologie) Herausgeber: Staatliches Museum für Naturkunde, Rosenstein 1, D-70191 Stuttgart Stuttgarter Beitr. Naturk. Nr. 694 | 64 5.,136 Abb. | Stuttgart, 20. IV. 2006 Revision of the Oriental species of the genus Bradymerus Perroud, with descriptions of 29 new species (Coleoptera: Tenebrionidae)! WOLFGANG SCHAWALLER Abstract The Oriental species of the genus Bradymerus Perroud (Coleoptera: Tenebrionidae: Cno- dalonini) are revised. The species characters are discussed and figured. —- New species: Brady- merus andoti n. sp. (Sulawesi), B. bocakorum n. sp. (Mindanao), B. crockerensis n. sp. (Borneo), B. emasicus n.sp. (Borneo), B. fouquei n.sp. (W Malaysia), B. gebieni n.sp. (Sumatra), B. gerstmeieri n. sp. (Sumatra), B. grimmi n.sp. (Borneo), B. hauseri n.sp. (W Malaysia), B. kan- chenjungicus n. sp. (Nepal), B. kaszabi n. sp. (Sulawesi), B. kinabalicus n.sp. (Borneo), B. ko- dadai n. sp. (Borneo), B. kulzeri n.sp. (Sulawesi), B. laoticus n.sp. (Laos), B. lombokicus n. sp. (Lombok), B. majeri n. sp. (Andaman Islands), B. malayicus n. sp. (W Malaysia, Thailand), 2. maramagicus n.sp. (Mindanao), B. masumotoi n. sp. (Island Lan Hsu near Taiwan), B. merk- lin.sp. (Vietnam), B. michihikoi n.sp. (Sulawesi), B. pseudomalayicus n.sp. (W Malaysia), B. reibnitzi n. sp. (Java), B. riedeli n. sp. (Sulawesi), B. sprecherae n. sp. (Sulawesi), B. sumatranus n.sp. (Sumatra), B. sumbawaicus n.sp. (Sumbawa), B. thailandicus n.sp. (Thailand). - New synonyms: B. clathratus Schaufuss, 1887 (B. junctus Shibata, 1980 n.syn.), B. sijthoffi Gebien, 1925 (B. tibialis Kulzer, 1951 n.syn.). - New combinations: Bradymerus aeneus (Kaszab, 1980) n. comb. from Planibates, Bradymerus fukiensis (Kaszab, 1954) n. comb. from Plani- bates, Derosphaerus opacicollis (Kulzer, 1951) n. comb. from Bradymerus. — Lectotypes are designated for: B. acutangulus Gebien, 1925, B. acuticostis Gebien, 1925, B. andamanus Ge- bien, 1914, B. aulacopterus (Fairmaire, 1883), B. celebensis Gebien, 1925, B. clathratus Schau- fuss, 1887, B. drescheri Gebien, 1925, B. eschscholtzi Gebien, 1921, B. impressicollis Gebien, 1913, B. interstitialis Schaufuss, 1887 (B. alternatus Schaufuss, 1887 syn.), B. mcgregori Ge- bien, 1921, B. nodicollis Gebien, 1925, B. pertyi Gebien, 1921 (B. elongatus Gebien, 1913 ho- monym), B. sijthoffi Gebien, 1925, B. spretus Gebien, 1925. Keywords: Coleoptera, Tenebrionidae, Cnodalonini, Bradymerus, Oriental, new species, new synonyms, taxonomy. ! Contributions to Tenebrionidae, no. 53. — For no. 52 see Annales Zoologici 55: 565-569 (2005). 2 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Zusammenfassung Die orientalischen Arten der Gattung Bradymerus Perroud (Coleoptera: Tenebrionidae: Cnodalonini) werden revidiert. Die Artkennzeichen werden diskutiert und abgebildet. - Neue Arten: Bradymerus andoi n.sp. (Sulawesi), B. bocakorum n.sp. (Mindanao), B. crocke- rensis n.sp. (Borneo), B. emasicus n.sp. (Borneo), B. fouquei n.sp. (W Malaysia), B. gebieni n.sp. (Sumatra), B. gerstmeierin.sp. (Sumatra), B. grimmin.sp. (Borneo), B. hauseri n.sp. (W Malaysia), B. kanchenjungicus n.sp. (Nepal), B. kaszabi n.sp. (Sulawesi), B. kinabalicus n.sp. (Borneo), B. kodadai n.sp. (Borneo), B. kulzeri n.sp. (Sulawesi), B. laoticus n.sp. (Laos), B. lombokicus n.sp. (Lombok), B. majeri n.sp. (Andaman Inseln), B. malayicus n.sp. (W Ma- laysia, Thailand), B. maramagicus n.sp. (Mindanao), B. masumotoi n. sp. (Insel Lan Hsu nahe Taiwan), B. merkli n.sp. (Vietnam), B. michihikoi n.sp. (Sulawesi), B. pseudomalayicus n. sp. (W Malaysia), B. reibnitzi n.sp. (Java), B. riedeli n.sp. (Sulawesi), B. sprecherae n.sp. (Sula- wesi), B. sumatranus n. sp. (Sumatra), B. sembawaicus n. sp. (Sumbawa), B. thailandicus n. sp. (Thailand). — Neue Synonyme: B. clathratus Schaufuss, 1887 (B. junctus Shibata, 1980 n.syn.), B. sijthoffi Gebien, 1925 (B. tibialis Kulzer, 1951 n.syn.). - Neue Kombinationen: Bradyme- rus aeneus (Kaszab, 1980) n. comb. von Planibates, Bradymerus fukiensis (Kaszab, 1954) n. comb. von Planibates, Derosphaerus opacicollis (Kulzer, 1951) n. comb. von Bradymerus. - Lectotypen werden designiert für: B. acutangulus Gebien, 1925, B. acuticostis Gebien, 1925, B. andamanus Gebien, 1914, B. aulacopterus (Fairmaire, 1883), B. celebensis Gebien, 1925, B. clathratus Schaufuss, 1887, B. drescheri Gebien, 1925, B. eschscholtzi Gebien, 1921, B. impres- sicollis Gebien, 1913, B. interstitialis Schaufuss, 1887 (B. alternatus Schaufuss, 1887 syn.), B. mcgregori Gebien, 1921, B. nodicollis Gebien, 1925, B. pertyi Gebien, 1921 (B. elongatus Ge- bien, 1913 Homonym), B. sijthoffi Gebien, 1925, B. spretus Gebien, 1925. Contents TEN OCLC OI Fe a ne en ee m A ne ne 8 ! DEESDEeIESIChATAEterS, put a a Ne ak EEE TEN. 5 3° Theknown Oriental’species of Bradymerus nn pad, Bea, Ped od dem 5 4 Descriptions of new Oriental species of Bradymerus .... 00. eee es 42 5 = Doubttultaxsof-Bradymerus N neu een de Dear ee BH Laie Ue thd eat hd epee 58 6, Transfer to.thegenusDeräsphaerus zn 8h 2 8 5 DIN BYE DORE Eee oh ack Dat BA, 59 7 Identification key for the Oriental species of Bradymerus ....... 000 c cece eee eee 59 B. References sake chee tla et, de ths ania dant Ain imeem mee a Sie Sete deen ace Mente 63 1 Introduction The numerous species of the genus Bradymerus Perroud, 1864 (type-species: tu- berculatus Perroud, 1864 = amicorum Fairmaire, 1849) are distributed mainly in the Indo-Australian Region, additional species are known from Micronesia, the Sey- chelles, Madagascar and the eastern Afrotropical Region. GEBIEN (1913, 1925) sum- marized the taxonomic knowledge of that time, subsequently Kurzer (1951), Ka- sZAB (1954, 1980) and SHiBATA (1980) added a few new Oriental species. In the present contribution, all species of Bradymerus are revised occuring in an area in southeastern Asia as indicated on the map (Fig. 1). Not included are speci- mens from the Moluccas (= Maluku Islands, for example Halmahera, Seram), these taxa are intended to be treated later together with the Papuan species. Also not in- cluded are the species from the Pacific Islands (Micronesia, see KULZER 1957) as well as the few ones from the Afrotropical area. This paper treats 44 known and adds 29 new species from the selected area. However, the total number of species in that re- gion is even higher, because some taxa remain doubtful (chapter 5) and some older specimens and some single females from different collections could not clearly be identified at present. The distributional data in this paper are not just cited from the SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 3 | we ee Se er a Fig. 1. The treated area in southeastern Asia. labels but are partly completed by additional data for a better localization, and trans- lated in several cases from other languages into English. To avoid superfluous repe- titions, the country names India and Indonesia are omitted in case of the Andaman Islands and Sunda Islands respectively. Borneo is used in a geographical and not po- litical sense (provinces Sabah and Sarawak belonging to Malaysia, Kalimantan be- longing to Indonesia). At present, the genus Bradymerus is classified into the tribe Cnodalonini, sub- family Stenochiinae (new names according to BOUCHARD et al. 2005). GEBIEN (1925) provided the diagnostic characters of the genus and already mentioned the difficul- ty of dividing the huge genus in natural species-groups. This is a general, still un- solved problem in tenebrionid taxonomy, occuring in many species-rich and widely distributed genera (for example in Amarygmus Dalmann, 1823, Gonocephalum Chevrolat, 1849, Laena Latreille, 1829, Strongylium Kirby, 1818, etc.). The used species-characters of adult Bradymerus are given in chapter 2, larval characters are nearly unknown. Not all features might be considered as ehe but are used traditionally in the present state of knowledge. Further studies under phylogenetical 4 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 aspects might show, that Bradymerus in the treated scope consists of paraphyletic or polyphyletic species assemblages. At least one species could be transferred herein from Bradymerus to Derosphaerus Thomson, 1858. On the other hand, it can also not be excluded, that further species, described in other genera, must be transferred to Bradymerus. In the collection of the Hungarian Natural History Museum, KaszaB labelled some species of Bradymerus under Calabosca Fairmaire, 1894, even when he had described two of these under Planibates Kaszab, 1939. Very probably, the genera Calabosca and Planibates are junior synonyms of Bradymerus — but for establishing this synonymy, the corresponding type species (Ascalabus pedi- noides Fairmaire, 1893 from Indochina for Calabosca and Planibates papuanus Kaszab, 1939 from New Guinea) should be reexamined during a phylogenetical analysis. The biology of the species of Bradymerus is nearly unknown. According to own observations, adult beetles are active during night in arboreal habitats, assemblages of several individuals often creeping around on old trees and rotten wood. Their lar- vae very probably live within those trees. The best method for collecting adult bee- tles is the search on old trees during night with an electric torch. Although all species possess fully developed wings, they are attracted by light only in single cases. In general, GEBIEN usually labelled the type material of his new species in German as “Type” and “Cotype”, but did not publish this in the corresponding descriptions. In other taxa he labelled the type specimens equally as “Type”. In all cases, in which the original descriptions are based on an unspecified number of syntypes (indepen- dent from the labelling), lectotypes and paralectotypes are designated herein in or- der to fix a single name-bearing type and thus to define the species, according to Ar- ticle 74.7.3 of the International Code of Zoological Nomenclature. Acronyms of depositories BMNH The Natural History Museum, London (Max BARCLAY) CBBB Collection Boris BUCHE, Berlin CKAO Collection Dr. Kryosu1 ANDO, Osaka CMLS Collection Martin LirLıc, Saarbrücken CRFL Collection RENE Fouquz, Liberec CRGT Collection Dr. ROLAND Grimm, Tübingen CRRH Collection ROLF Roper, Hasselby CRSW Collection RUDOLF SCHUH, Wien CSBC Collection STANISLAV BECvAR, Ceské Bud£jovice HNHM Hungarian Natural History Museum, Budapest (Dr. Orrö MERKL) MHNG Museum d’Histoire Naturelle, Genéve (Dr. GruLIO CuccoDoRo) MHNL Museum d’Histoire Naturelle, Lyon (Dr. HAROLD LABRIQUE) MNHUB Museum für Naturkunde der Humboldt-Universität, Berlin (BERND JÄGER) MNST Museum of Natural Science, Taichung (Dr. Kimo Masumoto) NHMB _ Naturhistorisches Museum, Basel (Dr. MicHEL BRANCUCCT) NHMB-F Naturhistorisches Museum, Basel, collection G. Frey (Dr. Eva SPRECHER) NHMC Natural History Museum and Institute, Chiba (Dr. Kınıo MAsumoOTo) NSMT National Science Museum, Tokyo (Dr. Kımıo Masumoto) SMNS Staatliches Museum für Naturkunde, Stuttgart TMSA Transvaal Museum, Pretoria (RUTH MULLER) ZFMK Zoologisches Forschungsinstitut und Museum ALEXANDER KOENIG, Bonn (Prof. Dr. M. SCHMITT) ZSM Zoologische Staatssammlung, München (Dr. Martin BAEHR) SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 5 Acknowledgements For the trustful and often extended loan of material I want to express first my gratitude to all friends and colleagues listed in the compilation of collections. The hospitality of Dr. Orro Mrrkı (Budapest), RUTH MÜLLER (Pretoria) and Dr. Eva SPRECHER (Basel) during my visits for comparative studies is greatly appreciated. Own field work in Asıa was supported during different trips by Dr. RoLAND Grimm (Tübingen), Dr. CHrIstoPH HÄUSER (Stuttgart), Dr. MartTIN Hauser (Urbana/lllinois), Prof. Dr. JOCHEN MARTENS (Mainz) and JÜRGEN TRAUT- NER (Filderstadt). Last but not least I thank JOHANNES REIBNITZ (Stuttgart) again for profes- sionally producing the photographs with the automontage system and for arranging the plates. 2 Species characters The combination of the following characters are considered as species-specific: Body length with a certain variability. Dorsal side ferrugineous, brownish or black- ish, without or with metallic shine. Genae distinctly broader or not broader than eyes. Frons with or without distinct supraorbital keels and/or supraorbital furrows. Last 3, 4, 5 or 6 antennomeres forming a club. Pronotum of different shape. Anteri- or corners of pronotum protruding or not protruding. Lateral margin of pronotum without or with distinct crenulation. Pronotal disc with rough and confluent or with fine and separate punctation, between punctures with or without granules. Pronotal disc with or without medial furrow/impression. Elytral intervals without or with tubercles/keels, equal on all intervals or only partly, often similar only on alternate intervals. Tibiae in both sexes externally with or without distinct keels. Sometimes sexually dimorphic femora and tibiae present in males (sometimes unvisible from dorsal and to be overlooked, so also by GEBIEN 1925). Aedeagus of different shape. Apart from the sexually dimorphic femora and tibiae in some species, the external characters of males and females are identical. Thus, a few single females are also de- scribed as new species if the external characters are adequate for taxonomic separa- tion — some further females must remain undescribed. 3 The known Oriental species of Bradymerus Bradymerus acutangulus Gebien, 1925 (Figs. 3, 74) Studied type material: Java, Preanger, leg. P. F. SyrHorr, 1 d syntype NHMB-F (la- belled by GEBIEN as type), designated herewith as lectotype. New material: None. Diagnostic characters: Dorsal view see Fig. 3, dorsal side dark ferrugineous without metallic shine, body length 6.5mm. Genae not broader than eyes, frons without distinct supraorbital keels but with deep supraorbital furrows. Last 5 an- tennomeres forming a club. Anterior corners of pronotum extremely protruding and acute, lateral margin without distinct crenulation, pronotal disc with rough punctation and without medial impression, between punctures without granules. Elytral intervals with high and partly confluent granules. Tibiae in both sexes exter- nally without distinct keels. Aedeagus see Fig. 74. Distribution: Java. 6 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Bradymerus acuticostis Gebien, 1925 (Figs. 2, 75) Studied type material: Philippines, Leyte, Tacloban, 1 d syntype NHMB-F (labelled by GEBIEN as type), designated herewith as lectotype. New material: Philippines, Leyte, leg. BOETTCHER, 1 ex. BMNH (acuticollis sic! det. Kurzer). — Philippines, Leyte, Lake Danao, 500m, 19.11.1991, leg. W. SCHAWALLER, 4 ex. SMNS. - Philippines, Leyte, Visca N Baybay, 28.11.1991, leg. W. SCHAWALLER, 1 ex. SMNS. - Philippines, Samar, 3 ex. MNHUB. - Philippines, Mindanao, Misamis, Don Victoriano, 1700 m, 1.-3.V.1996, leg. L. Borm, 1 ex. SMNS. - Philippines, SE Luzon, Vivac, leg. BOETTCHER, 1 ex. HNHM. - Philippines, Mindanao, Surigao, leg. C. F. BAKER, 1 ex. HNHM. — Philippines, Romblon Prov., Tablas, St. Augustin, Dubduban, Busai Falls, 23.-26.X1.1994, leg. H. ZETTEL, 1 ex. HNHM. - Micronesia, Palau Islands, 2002-2003, leg. K. TAKAHASHI, 17 ex. NSMT, 3 ex. SMNS. Diagnostic characters: Dorsal view see Fig. 2, dorsal side dark ferrugineous without metallic shine, body length 5.3-6.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, prono- tal disc with rough punctation and with a longitudinal impression, between punc- tures without granules. All elytral intervals with distinct and similar keels, keels with granules. Tibiae in both sexes externally without keels; in males posterior tibia inter- nally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 75. Distribution: Philippines (Luzon, Leyte, Samar, Mindanao), Micronesia (Palau Islands). Bradymerus aeneus (Kaszab, 1980) n. comb. (Fig. 4) Planibates aeneus Kaszab, 1980. Studied type material: Vietnam, Nghe tinh, NW Quy chau (locality labelled in kyril- lic letters), 500m, 11.1I1.1962, leg. KABakov, 1? paratype of Planibates aeneus HNHM (labelled subsequently by Kaszas as Calabosca aenea). New material: None. Diagnostic characters: Dorsal view see Fig. 4, dorsal side dark ferrugineous with a feeble metallic shine, body length 8.3 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 3 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with separate punctation and with distinct medial impression, between punctures without granules. All elytral intervals slightly convex and without keels or granules. Tibiae without distinct keels. Aedeagus unknown, only female available. Remarks: Bradymerus aeneus is placed by Kaszas in Planibates Kaszab, 1939, although he had no males available (the holotype and the single paratype are fe- males). The modified anterior tibia in males is said to be a diagnostic character for Planibates. However, this character is not generic, all other characters coincide with Bradymerus. The assignment of this species to different genera by Kaszag demon- strates once more the confusion about the genera Bradymerus, Planibates and Cala- bosca. See also the remarks in the introduction and under Bradymerus fukiensis. Distribution: Vietnam. Bradymerus aeratus Gebien, 1925 (Fig. 5) Studied type material: Sri Lanka (labelled as Ceylon), 2 holotype NHMB-F (holo- type by monotypy, labelled by GEBIEN as type). New material: None. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 7 Diagnostic characters: Dorsal view see Fig.5, dorsal side dark ferrugineus, with distinct metallic shine, body length 10.0mm. Genae not broader than eyes, frons without distinct supraorbital keels but with deep supraorbital furrows. Last 5 antennomeres forming a club. Anterior corners of pronotum protruding and acute, lateral margin without distinct crenulation, pronotal disc with confluent punctation and without medial impression, between punctures without granules. Internal ely- tral intervals slightly convex and with a few indistinct granules, intervals 3, 4 poste- riorly and intervals 5-8 completely with keels, keels without granules. Tibiae exter- nally without distinct keels. Aedeagus unknown, only female available. Distribution: Sri Lanka. Bradymerus alternicostis Gebien, 1913 (Figs. 6, 76) Studied type material: Philippines, Luzon, Laguna, Mt. Banahao, leg. C. S. Banks, 3 holotype NHMB-F (holotype by monotypy, labelled by GEBIEN as type). New material: None. Diagnostic characters: Dorsal view see Fig. 6, dorsal side dark ferrugineous without metallic shine, body length 8.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and with a feeble medial impression, be- tween punctures without granules. Alternate elytral intervals 3, 5, 7 with keels, in- tervals 2, 4, 6 smooth and without punctures and granules, keels without granules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 76. Distribution: Philippines (Luzon). Bradymerus andamanus Gebien, 1914 (Figs. 7, 77) Studied type material: Andaman Islands, 1 ? syntype NHMB-F (labelled by GEBIEN as type), designated herewith as lectotype. New material: Andaman Islands, leg. DE ROEPSTORFF, 2 ex. MNHUB (det. GeBIEN, 1, 1 ex. without abdomen). — Andaman Islands (labelled as Andamanen), 1 d SMNS. Diagnostic characters: Dorsal view see Fig. 7, dorsal side ferrugineous with- out metallic shine, body length 7.0-8.5mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 4 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and with medial furrow, between punctures without granules. Elytral intervals slightly convex and without keels or granules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 77. Remarks: See remarks under Bradymerus atronitens. Distribution: Andaman Islands. Bradymerus aratus Fairmaire, 1896 (Figs. 9, 79) Bradymerus interruptus Gebien, 1914 syn. Studied type material: Borneo, Njabang, d holotype of B. interruptus NHMB-F (holotype by monotypy, labelled by GEBIEN as type). New material: Sumatra, coll. Hauser, 1 ex. MNHUB (labelled by FAIRMAIRE as Bradymerus aratus n.sp., non-type specimen because not listed in the description). — Sumatra, Fort de Kock, 920m, 1925, leg. E. Jacogson, 1 ex. BMNH (aratus det. GEBIEN). — Borneo, Sabah, Crocker Range NP, NW Keningau, 900-1200 m, 16.-20.X1.1996, leg. W. SCHAWALLER, 5 ex. SMNS. - Borneo, Sabah, Kundasang, 1450 m, 19.V.2005, leg. R. Grimm, 1 ex. CRGT. - 8 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Borneo, Sabah, Crocker Range, Gunung Emas, 15.-27.1V.1993, leg. I. JEnı$ & M. Stra, 2 ex. ZSM. — Borneo, Brunei, 1 ex. HNHM. - Borneo, S Kalimantan, Loksado, 1000 m, 7.-22.1X.1997, leg. S. JAK, 1 ex. CSBC. — W Malaysia, 90 km NE Ipoh, Banjaran Titi Wangsa, Mt. Gerah, 1900 m, 1.-17.IV.2000, leg. P. CecHovsky, 1 ex. ZSM. Diagnostic characters: Dorsal view see Fig. 9, dorsal side ferrugineous with- out metallic shine, body length 4.5-6.0 mm. Genae broader than eyes, frons with dis- tinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and with medial impression, between punctures with distinct granules. Internal elytral intervals smooth with small granules, external intervals with high longitudinal granules. Tibiae in both sexes externally with distinct keels. Aedeagus see Fig. 79. Distribution: Sumatra (type locality of aratus), Borneo (type locality of inter- ruptus); W Malaysia (new record). Bradymerus asper Kulzer, 1951 (Figs. 11, 80) Studied type material: Philippines, Mindanao, Butuan, leg. C. F Baker, 2 holotype NHMB-E. New material: Philippines, Luzon, Nueva Viscaya, Imugan (labelled as Imugin), leg. C. FE. Baker, 1 6 BMNH. Diagnostic characters: Dorsal view see Fig. 11, dorsal side ferrugineous with- out metallic shine, body length 4.5-5.5 mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and with medial impression, between punctures with granules. Elytral intervals with distinct longitudinal granules, on intervals 3 and 5 partly con- fluent and forming keels, keels with granules. Tibiae in both sexes externally with distinct keels. Aedeagus see Fig. 80. Distribution: Philippines (Mindanao, Luzon). Bradymerus atronitens Kulzer, 1951 (Figs. 10, 83) Studied type material: Philippines, Luzon, Imugan, leg. BOETTCHER, d holotype NHMB-F (in bad condition, both antennae and all right legs missing). — Philippines, Luzon, Baguio, 1 paratype HNHM (labelled by Kaszag as Calabosca atronitens). New material: Philippines, Luzon, Nueva Viscaya, Imugan (labelled as Imugin), leg. C. F. Baker, 1 ex. BMNH. - Philippines, Luzon, Baguio, 1 ex. SMNS (non-type). — Central Laos, Khammouan Prov., Ban Khoun Ngeun, 200 m, 24.-29.IV. and 19.-31.2001, leg. L. DEm- BICKY, 7 ex. SMNS. Diagnostic characters: Dorsal view see Fig. 10, dorsal side blackish without metallic shine, body length 8.0-9.2 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 4 antennomeres forming a club (in non-type mater- ial and from the description). Anterior corners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with separate punctation and with feeble medial impression, between punctures without granules. Elytral inter- vals slightly convex and without keels or granules. Tibiae in both sexes externally without distinct keels; in males anterior tibia internally in the distal third with an in- distinct (holotype) or quite distinct tooth (non-type material from Laos). Aedeagus see Fig. 83. Remarks: The anterior tibia in the male holotype bears internally only an indis- SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 9 tinct tooth, in the new material from Laos this tooth is distinctly developed. All oth- er external characters coincide between the holotype from the Philippines and the new material from Laos, also the quite extraordinary shape of the parameres. Quite similar in external characters (with completely unarmed anterior male tibia) as well as in the shape of the aedeagus is Bradymerus andamanus from the Andaman Is- lands. Further material from different localities might show, that B. atronitens is a ju- nior synonym of B. andamanus. Distribution: Philippines (type locality Luzon); Laos (new record). Bradymerus aulacopterus (Fairmaire, 1883) (Figs. 12, 82) Bradynocerus aulacopterus Fairmaire, 1883. Studied type material: Selayar Island (labelled as Saleyer Island), collection R. OBERTHUR, 1d syntype ZFMK (labelled as holotype, but not by the author), designated herewith as lectotype. New material: None. Diagnostic characters: Dorsal view see Fig. 12, dorsal side blackish without metallic shine, body length 10.8mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with separate punctation and without impression, between punctures without gran- ules. Elytral intervals slightly convex, alternate intervals 3, 5, 7 posteriorly with fee- ble keels, all intervals without granules. Tibiae externally without distinct keels; in males posterior tibia internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 82. Remarks: The genus Bradynocerus Fairmaire, 1883 is said to be related to Tetra- phyllus and Camaria, but the synonymy with Bradymerus is doubtless and already listed by GEBIEN (1936-1944) in his catalogue. See also Bradymerus masumotoi n. sp. from the island Lan Hsu near Taiwan. Distribution: Selayar Island (S Sulawesi). Bradymerus bifurcatus Kaszab, 1980 (Figs. 13, 81) Studied type material: Vietnam, Bac Thai, Mts. 50km NE Thai Nguyen, 300m, 10.1V.1963, leg. O. KABAKov, 1 d paratype HNHM. New material: N Vietnam, Quang Ninh, Ha Long, 29.V.-1.V1.1985, leg. A. OLEXa, 1 ex. SMNS. - S Thailand, Betong, Yala Distr., Gunung Cang Dun, 25.111.-22.1V.1993, leg. J. HorAk, 1 ex. CSBC. Diagnostic characters: Dorsal view see Fig. 13, dorsal side dark ferrugineous without metallic shine, body length 7.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels but with deep supraorbital furrows. Last 5 an- tennomeres forming a club. Anterior corners of pronotum protruding, lateral mar- gin with feeble crenulation, pronotal disc with confluent punctation and without medial impression, between punctures without granules. Alternate elytral intervals 3 (only anteriorly), 5, 7 with keels, intervals 2, 4, 6 flat and without granules, keels with granules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 81. Distribution: Vietnam (type locality), Thailand (new record). 10 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Bradymerus caeruleipennis Gebien, 1913 (Fig. 15) Studied type material: Philippines, Luzon, Benguet, Mt. Pulog, leg. H. M. Curran, ? holotype NHMB-F (holotype by monotypy, labelled by GEBIEN as type). New material: None. Diagnostic characters: Dorsal view see Fig. 15, dorsal side blackish, with dis- tinct metallic shine, body length 10.0 mm. Genae not broader than eyes, frons with- out distinct supraorbital keels. Last 4 antennomeres forming a club. Anterior cor- ners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with fine and separate punctation and without medial impression, be- tween punctures without granules. Elytral intervals slightly convex and without keels or granules. Tibiae externally without distinct keels. Aedeagus unknown, only female available. Distribution: Philippines (Luzon). Bradymerus carinatus Fairmaire, 1886 (Figs. 19, 85) Bradymerus corinthius Fairmaire, 1896 syn. Studied type material: None. New material: Philippines, Luzon, 3 ex. MNHUB (det. Gesien, Hist. Coll., nos. 46463-5). — Philippines, Luzon, Mt. Banahao, 14.VIII.1914, leg. BOETTCHER, 3 ex. MNHUB. - Philippines, Leyte, 3 ex. MNHUB. - Philippines, Luzon, Balbatan, leg. BOETTCHER, 1 ex. NHMB-F (det. Gegen). — Philippines, Vian, 1 ex. HNHM. - Philippines, Mindanao, coll. J. THomson, 1 ex. HNHM. - Philippines, Mindanao, Surigao, leg. C. E Baker, 1 ex. ZSM. — Philippines, Manila, leg. Bates, 1 ex. ZSM. - Philippines, N Luzon, Kalinga-Apayao, Cordillera Central, Saltan Valley, 750 m, 22.111.2000, leg. L. DEmBicky, 4 ex. SMNS. - Philippines, N Luzon, Kalinga-Apayao/Abra Province Boundery, Cordillera Cen- tral, Pass, 1500-1700 m, 26.-28.111.2000, leg. L. DEMBICKY, 1 ex. SMNS. — Philippines, Panay, Cambunao Iloilo, Mt. Tinagung, 300-1000 m, V.1994, leg. L. MoHacan, 1 ex. HNHM. Diagnostic characters: Dorsal view see Fig. 19, dorsal side blackish, with dis- tinct metallic shine, body length 10.0-10.3 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, prono- tal disc with dense punctation and without medial impression, between punctures without granules. Elytral intervals slightly convex, external intervals with feeble keels, keels smooth without granules. Tibiae in both sexes externally without dis- tinct keels. Aedeagus see Fig. 85. Distribution: Philippines (Luzon, Leyte, Mindanao, Panay). Bradymerus celebensis Gebien, 1925 (Figs. 20, 86) Studied type material: S Celebes (Sulawesi), coll. ScHauFuss, 1 d syntype MNHUB (labelled by GEBIEN as type), designated herewith as lectotype. New material: N Sumatra, Brastagi, 900 m, 20.VII.1980, leg. E. Hess, 3 ex. HNHM, 1 ex. SMNS. — N Sumatra, Brastagi, Gunung Sibayak, 1450-1900 m, 19.-23.11.1991, leg. L. Bocdk & M. BocAxova, 12 SMNS. — N Sumatra, Brastagi, 28.11.1997, leg. N. Kanıe, 1 ex. CKAO. Diagnostic characters: Dorsal view see Fig. 20, dorsal side ferrugineous with- out metallic shine, body length 7.5-8.0 mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and with medial impression, between punctures with granules. Al- ternate elytral intervals 1 (only posteriorly), 3, 5, 7 with keels, keels with granules, SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 11 intervals 2, 4, 6 without granules. Tibiae in both sexes externally with distinct keels; in males posterior tibia internally near the tip with a row of distinct long, yellow se- tae. Aedeagus see Fig. 86. Remarks: In the specimens from Sumatra, the elytral interval 1 along the suture bears a more prominent keel, in the lectotype from Sulawesi this keel exists only in the posterior part, all other diagnostic characters coincide. Thus I hope not to fail in assigning the zoogeographically remarkable new findings from Sumatra to this species. It is striking that I could not find this species among numerous material of Bradymerus from Sulawesi, thus the lectotype might be erroneously labelled. Bradymerus serricollis from Sri Lanka and Yunnan is quite similar, but can be sepa- rated by a smaller body size in the average (5.5—7.0 mm), by broad scale-like setae on pronotum and elytra (thin acute setae in B. celebensis) and by unmodified posterior tibiae in males. The aedeagi of both species are similar (Figs. 86, 132). Distribution: Sulawesi (type locality); Sumatra (new record). Bradymerus clathratus Schaufuss, 1887 (Figs. 16, 87) Bradymerus aequecostatus Fairmaire, 1893 syn. Bradymerus junctus Shibata, 1980 n. syn. Studied type material: Without exact data, 1 2 syntype of B. clathratus MNHUB, des- ignated herewith as lectotype. — S Celebes (Sulawesi), Bantimurang, 1882, leg. Ripe, 1 d ex. MNHUB, might be a syntype of B. clathratus (JAEGER in litt.). New material: Taiwan, Shanmei, 600 m, 23.V.1977, leg. J. KLAPPERICH, 1 ex. SMNS (det. Kaszas). — Taiwan, Hoozan, 1.1910, leg. H. SAUTER, 5 ex. MNHUB. - Taiwan, Fuhosho, IX.1909, leg. H. SAUTER, 5 ex. MNHUB. - Taiwan, Kangding, 1992-1993, leg. K. Masumo- TO, 41 ex. NSMT. — Taiwan, Nantou, 11.1V.1998, leg. Lien Yu, 1 ex. CKAO. - Taiwan, Kent- ing Park, 14.V1.1971, leg. Y. Marpa, 1 ex. CKAO. - Tatwan, Ho Shan, 30.V.1995, leg. K. AN- DO, 7 ex. CKAO, 3 ex. SMNS. — Taiwan, Island Lan Hsu, 5.-8.V1.1986, leg. S. Osawa, 2 ex. NSMT. - Taiwan, Island Lanyu (= Lan Hsu), 24.1V.1998, leg. K. Masumoto, 1 ex. NSMT. - Taiwan, Island Lanyu (= Lan Hsu), 10.X.1970, leg. Y. Krvoyama, 4 ex. CKAO. - Japan, Ryukyu Islands, Ishigaki Island, III.-VIIL.1968, leg. O. Sato, 2 ex. CKAO. - Japan, Ryukyu Islands, Hateruma Island, 27.VII.1964, leg. M. Yasut, 2 ex. CKAO. - Japan, Okinawa Pref., Iriomote Island, Ohara forest, 19.IX.2003, leg. P. JALoszyNsk1, 3 ex. HNHM. — Andaman Is- lands, leg. DE ROEPSTORFF, 5 ex. MNHUB. - Andaman Islands, Havelock Island, village no. 7, 22.1V.-14.V.1998, leg. K. & S. Mayer, 7 ex. NHMB, 2 ex. SMNS. - India, Upper Assam, leg. S. HARTERT, 3 ex. MNHUB. - Thailand, Chumphon Prov., Pha To, 27.1. -14.1V.1996, leg. K. Majer, 2 ex. SMNS. - S Thailand, Satun Prov., Thale Ban, 200 m, 8.-13.1V.1997, leg. J. KouigAc, 4 ex. NHMB. - S Thailand, Betong, 23.-24.1V.1992, leg. J. HorAx, 10 ex. ZSM, 1 ex. CRGT. - E Thailand, Chanthaburi Distr., Khao Soi Dao, 5.-13.V.1998, leg. J. HORAR, 3 ex. CRGT. - Thailand, Phuket, Kamala, 12.11.2002, leg. P. Dynorr, 1 ex. SMNS. - Thailand, Trat Prov., Ko Chang Island, 0-200 m, 1.-5.VII.2004, leg. R. & H. Fouqu#, 31 ex. CRFL. - W Malaysia, Pahang, Tioman Island, Kg. Terek, 15.—26.VII.1992, leg. R. SCHUH, 1 ex. CRSW. — W Malaysia, Perak, 30km SE Ipoh, Cameron Highlands, Ringlet, 25.1V.-5.V.2001, leg. P. CecHovsky, 1 ex SMNS. - W Malaysia, Cameron Highlands, 2.I11.1974, leg. Y. Kıyovama, 1 ex. CKAO.- W Malaysia, Perak, 25 km NE Ipoh, Banjaran Titi Wangsa Mts., Mt. Korbu, 1200m, 1.-15.IV.2000, leg. P. CecHovsxkY, 3 ex. SMNS. — W Malaysia, Benom Mts., 15km E Kampong Dong, 700 m, 1.1V.1998, leg. L. DemBicxy & P. PACHOLATKO, 2 ex. NHMB. - W Malaysia, Cameron Highlands, 17.V.1981, leg. T. Harayama, 1 ex. NSMT. - W Malaysia, Kelantan, Gua Musang, 24.-28.VII.2001, leg. R. Fouquk & H. BArLOVA, 1 ex. CRFL. - W Malaysia, Terengganu, Kapong Bintang, between K. Baharu and K. Terengganu, 16.-19.VII.2001, leg. R. Fouquz & H. BARLOVA, 19 ex. CRFL. - Borneo, Kalimantan, Bu- sang/Rekut confluence, VIII.2001, leg. BRENDELL & MENDEL, 1 ex. BMNH. - Borneo, W Kalimantan, Nangah Serawai Distr., Tontang, 24.-30.VII.1993, leg. R. DuNpDaA & J. SCHNEIDER, 2 ex. CRFL. - Borneo, Sabah, Bunsit, 25.V.1988, 32 ex. NSMT. - Borneo, Sabah, 31km N Tenom, 3.V.1999, leg. M. SNtZEK, 2 ex. CMLS. — Borneo, Sabah, Tawau Hills Park, 12 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Tawau River, 8.V1.1998, leg. J. Kopapa, 1 ex. SMNS. - Borneo, Sabah, Kampung Takala, Kin- abatangan River, 5.V1.1998, leg. J. Kopapa, 2 ex. SMNS. — Borneo, Sabah, Batu Punggul Re- sort, 24.VI.—1.VII.1996, leg. J. Kopapa, 2 ex. SMNS. - Borneo, Sabah, 15 km S Tenom, 450 m, 11.V.2005, leg. R. Grimm, 32 ex. CRGT. - Borneo, Sabah, Tambunan, 600 m, 9.V.2005, leg. R. Grim, 24 ex. CRGT. - Borneo, Sabah, Keningau, 10.-20.X.1988, leg. M. Iron, 6 ex. CKAO. — Borneo, Sabah, Crocker Range NP, NW Keningau, 900-1200 m, 16.-20.X1.1996, leg. W. SCHAWALLER, 3 ex. SMNS. — Borneo, Sabah, Poring Hot Springs, 500 m, 29.X1.-2.X11.1996, leg. W. ScHAWALLER, 1 ex. SMNS. - Borneo, Sabah, Poring Hot Springs, 485m, 14.-31.VII1.1988, leg. A. SMETANA, 1 ex. MHNG. - Borneo, Sarawak, Belaga, Long Linau, 17.-21.111.1990, leg. A. Rıeper, 4 ex. SMNS. — Borneo, W Sarawak, Quop, II.-V.1914, leg. G. E. Bryant, 10 ex. BMNH. — W Sumatra, Siberut Island, IX.1924, leg. C. B. K. & N.S., 1 ex. BMNH (clathratus det. BLatr). - W Sumatra, Payakumbuh, Harau Valley, 1000 m, 9.-29.X.1991, leg. A. Rıeper, 1 ex. SMNS. -— W Sumatra, Bengkhulu, S Muko Muko, 16.VIII.1991, leg. D. ErBER, 4 ex. SMNS. — E Sumatra, Riau Prov., Bukit Tigapuluh NP, 18.-25.1.2000, leg. J. BEZDER, 1 ex. SMNS. — Lombok, Senaro, N slope of Rinjani, 1100 m, 2.-5.11.1994, leg. L. Borm, 10 ex. SMNS. - Java, Mt. Kawi, 1 ex. HNHM. - Java, Trawas, Gu- nung Penanggungan, E slope, 1000 m, 6.-9.V.2001, leg. L. Borm, 2 ex. NHMB. - W Java, Bo- gor, Kebun Raja, 13.1X.2005, leg. B. BUCHE, 8 ex. CBBB. — W Bali, Mt. Bahukaru, 1100 m, 26.X.2005, leg. E. Heiss, 1 ex. CRGT. — Sulawesi, Poso Distr., Tentena to Bada, 20.1V.-2.V.1994, leg. M. HIERMEIER, 1 ex. CRGT. - SE Sulawesi, Island Buton-Wakarumba, 3.-7.11.1994, leg. I. JEnı$ & M. Srrpa, 6 ex. ZSM. - Central Sulawesi, W Coast of Lake Poso, Taipa, 10.-11.1V.1999, leg. S. BECvAR & P. ZABRANSKY, 3 ex. CSBC. - Central Sulawesi, 20-35km NW Palopo, 1000-1400 m, 4.-5.IV.1999, leg. S. BECvAR & P. ZABRANSKY, 1 ex. CSBC. - Philippines, Luzon, Camarines Sur, 12km N Ocampo, 800 m, 25.-27.V.1996, leg. L. Borm, 11 ex. SMNS. - Philippines, Palawan, Port Barton, 150m, 14.-18.XII.1990, leg. L. Boum, 1 ex. NHMB. - Philippines, Palawan, Talabigan Barrio, 24.111.1979, leg. K. Wapa, 1 ex. NSMT. - Philippines, Island Basilan near Mindanao, leg. BOETTCHER, 1 ex. MNHUB. - Philippines, Island Sibuyan, Romblon, 1 ex. CKAO. - Philippines, Island Panay, Cambunao Iloilo, Mt. Tinagung, 300-1000 m, V.1994, leg. L. MoHAGAN, 1 ex. HNHM. Diagnostic characters: Dorsal view see Fig. 16, dorsal side dark ferrugineous without metallic shine, body length 5.0-7.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, prono- tal disc with rough punctation and with medial impression, between punctures with- out granules. Internal elytral intervals with weak, external intervals with distinct keels, keels with granules. Tibiae in both sexes externally without distinct keels; in males posterior tibia internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 87. Synonymy: SHiBaTa (1980) described Bradymerus junctus from the small Island Lan Hsu (= Lanyu, = Orchid Island) southeast of Tatwan and compared this taxon with B. acuticostis from the Philippines (but not with B. clathratus common on Tai- wan). The newly collected specimens from this island listed above fully coincide with specimens of B. clathratus from Taiwan and also with the original description of B. junctus including the therein given photograph. The posterior male tibia of B. junctus is said to have internally a dense row of yellow hairs, as present in B. clathra- tus. Therefore B. junctus Shibata, 1980 is considered as a new junior synonym of B. clathratus Schaufuss, 1887, although the type of B. junctus could not be examined. Remarks: I have seen specimens from the island Mauritius in the Indian Ocean (2 ex. TMSA, 1 ex. SMNS), which probably belong to Bradymerus clathratus. But without a reexamination of all African and Madagascaran taxa this identification is tentative. Distribution: Widespread in the Oriental region, known from Thailand, W Malaysia, India including Andaman Islands, Japan (Ryukyu Islands), Taiwan in- SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 13 cluding the adjacent Island Lan Hsu, Sunda Islands, Sulawesi and the Philippines (Luzon, Palawan, Panay, Basilan, Sıbuyan). Bradymerus crassicollis Kulzer, 1951 (Fig. 17) Studied type material: Philippines, Mindanao, Davao, leg. C. F BAKER (not BoETT- CHER!), 2 holotype NHMB-F. New material: None. Diagnostic characters: Dorsal view see Fig. 17, dorsal side blackish, with dis- tinct metallic shine, body length 11.5 mm. Genae not broader than eyes, frons with- out distinct supraorbital keels. Last 4 antennomeres forming a club. Anterior cor- ners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and without medial impression, between punctures without granules. Elytral intervals 3 (posteriorly), 5, 6, 7 with keels, keels without granules. Tibiae externally without distinct keels. Aedeagus unknown, only female available. Distribution: Philippines (Mindanao). Bradymerus crenulicollis Fairmaire, 1882 (Figs. 18, 91) Bradymerus denticeps Gebien, 1914 syn. Studied type material: None. New material: W Malaysia, Perak, Bukit Larut (Maxwell Hill), Taiping to Bintang Mts., 1100-1440 m, 9.-13.VII.2001, leg. R. Fouquz & H. BARLOVA, 6 ex. CRFL. —- W Malaysia, Terengganu, Kapong Bintang, between K. Baharu and K. Terengganu, 16.-19.VII.2001, leg. R. Fouqué & H. BARLOVA, 1 ex. CRFL. — Andaman Islands, Havelock Island, village no. 7, 22.1V.-14.V.1998, leg. K. & S. Majer, 4 ex. NHMB, 2 ex. SMNS. — W Sumatra, Payakumbuh, Harau Valley, 9.-29.X.1991, leg. A. RIEDEL, 1 ex. SMNS. — W Sumatra, Bengkhulu, 20km S Muko Muko, 16.VIII.1991, leg. D. ErBEr, 3 ex. SMNS. - W Sumatra, Bengkhulu Prov., Cu- rup, Bukit Kaba Mt., 1000-1500 m, 30.1.-3.11.2000, leg. J. BEZDER, 1 ex. SMNS. — W Sumatra, Pelompek (Kerinci), 13.-15.V.1991, leg. J. Moravec, 8 ex. NHMB, 2 ex. SMNS. - Sumatra, Aceh, Belankejeren, 1000 m, 10.X.1991, leg. W. BARRIEs, 3 ex. CRSW. - S Sumatra, Lampung Prov., Bukit Barisan Selatan NP, 5km SW Liwa, 600m, 7.-17.11.2000, leg. J. BEZDER, 1 ex. SMNS. - Sumatra, 1 ex. MNHUB (det. FAIRMAIRE). - W Sumatra, Baso, 800 m, III.1926, leg. E. Jacogson, 1 ex. BMNH (spretus det. GEBIEN). — SE Borneo, leg. S. V. GRABOwsKY, 1 ex. MNHUB. - Borneo, Sabah, Poring Hot Springs, 485 m, 19.VIII.1988, leg. A. SMETANA, 1 ex. MHNG. - Borneo, Sabah, Poring Hot Springs, 15.-30.X1I.1995, leg. C. HÄuser, 1 ex. SMNS. — Borneo, Sabah, Crocker Range NP, NW Keningau, 900-1200 m, 16.-20.X1.1996, leg. W. SCHAWALLER, 1 ex. SMNS. - Borneo, Sabah, 48km NW Ranau, 800 m, 20.V.2005, leg. R. GRIMM, 3 ex. CRGT. - Borneo, Sabah, Kimanis road near Keningau, 7.TV.1994, leg. N. KANIE, 1 ex. CKAO. - Borneo, Sabah, Crocker Range, Gunung Emas, 15.-27.IV.1993, leg. I. JENS & M. Srrsa, 3 ex. ZSM. - Borneo, Sarawak, Kapit Distr., Sebong, Baleh River, 9.-21.111.1994, leg. J. HorAx, 1 ex. ZSM. - Borneo, Sarawak, 19.-26.111.1995, leg. M. Iron, 3 ex. CKAO. - Central Sulawesi, 17 km E Pendolo, 800 m, 4.-9.VII.1999, leg. L. Bom, 1 ex. SMNS. - Java, Preanger, Mt. Tangkoeban, Prahoe, 1800 m, XI.1935, 1 ex. NHMB-F (det. Kurzer). — Java, Mt. Kawi, 3 ex. HNHM. - Java, leg. DE Haan, 1 ex. MNHUB (Hist. Coll. no. 47989). — W Ja- va, Bogor, Kebun Raja, 18.1X.2005, leg. B. Büchte, 4 ex. CBBB. Diagnostic characters: Dorsal view see Fig. 18, dorsal side ferrugineous with- out metallic shine, body length 4.7-6.0 mm. Genae broader than eyes, frons with dis- tinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and without medial impression, between punctures with granules. Elytral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct 14 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 keels, keels with granules. Tibiae in both sexes externally with distinct keels. Aedea- gus see Fig. 91. Remarks: The published record from the Philippines (Mindanao, Iligan) by GE- BIEN (1925) very probably refers to the quite similar Bradymerus difficilis known from the same locality. Distribution: Sunda Islands, Sulawesi; W Malaysia, Andaman Islands (both new records). Bradymerus cucullatus Fairmaire, 1897 (Fig. 23) Studied type material: None. New material: India, Mumbai, Matheran, 800m, 1902, leg. Biro, 1 2 HNHM (det. Kaszas). Diagnostic characters: Dorsal view see Fig. 23, dorsal side ferrugineous with- out metallic shine, body length 4.8mm. Genae not broader than eyes but clypeus distinctly bent upwards, frons without distinct supraorbital keels. Last 5 anten- nomeres forming a club. Anterior corners of pronotum not protruding, but anterior margin medially with striking prolongation covering the head completely, lateral margin without distinct crenulation, pronotal disc with rough punctation and with- out medial impression, between punctures without granules. All elytral intervals slightly convex and with a row of fine granules. Tibiae externally without distinct keels. Aedeagus unknown, only female available. Distribution: India. Bradymerus cyaneipennis Pascoe, 1883 (Fig. 24) Studied type material: None. New material: Sri Lanka, 1910, leg. G. Lewis, 1 2 NHMB-F (compared with the type by Barr). Diagnostic characters: Dorsal view see Fig. 24, dorsal side blackish, with dis- tinct metallic shine, body length 11.5 mm. Genae not broader than eyes, frons with- out distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior cor- ners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with fine separate punctation and without medial impression, between punctures without granules. Elytral intervals slightly convex and without keels or granules. Tibiae externally without distinct keels. Aedeagus unknown, only female available. Distribution: Sri Lanka. Bradymerus difficilis Gebien, 1925 (Figs. 21, 90) Studied type material: Philippines, Mindanao, Iligan, leg. C. F. Baker, d holotype NHMB-F (holotype by monotypy, labelled by GEBIEN as type). New material: Philippines, Mindanao, 30 km E Malaybalay, Busdi, 1000 m, 5.-9.V.1996, leg. L. Bot, 6 ex. SMNS. - Philippines, Mindanao, Momungan, 1 ex. HNHM. - Philippines, Leyte, leg. BOETTCHER, 1 ex. HNHM. Diagnostic characters: Dorsal view see Fig. 21, dorsal side ferrugineous with- out metallic shine, body length 5.3-6.0 mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and without medial impression, between punctures with granules. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 15 Elytral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct keels, keels with granules. Tibiae in both sexes externally with distinct keels. Aedea- gus see Fig. 90. Remarks: Very similar to Bradymerus crenulicollis from the Sunda Idslands and B. serricollis from Sri Lanka, but, apart from a different shape of the aedeagus, the se- tae on the elytra are thin and acute in B. difficilis. Distribution: Philippines (Leyte, Mindanao). Bradymerus drescheri Gebien, 1925 (Figs. 26, 89) Studied type material: Java, Semareang, V1.1896, leg. DRESCHER, 1 d syntype (labelled by GEBIEN as type), designated herewith as lectotype. New material: Java, G. Kawi, VII.1934, leg. v. DorsBurc, 1% HNHM (labelled by Kaszag as Calabosca sp.). — Java, Malang, 1 ¢ SMNS. - Java, leg. F. Bates, 3 ex. BMNH (drescheri det. BLAIR). Diagnostic characters: Dorsal view see Fig. 26, dorsal side ferrugineous with- out metallic shine, body length 7.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 4 antennomeres forming a club. Anterior corners of pronotum slightly protruding, lateral margin without distinct crenulation but sinu- ated, pronotal disc with dense punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3, 5 with feeble keels, interval 7 with a more distinct keel, intervals 2, 4, 6 slightly convex, without granules. Tibiae in both sexes externally without distinct keels; in males anterior tibia internally in the distal third with a distinct tooth. Aedeagus see Fig. 89. Remarks: The sexually dimorphic male tibia of Bradymerus drescheri was not mentioned in the original description by GEBIEN (1925). The antenna has a 4-seg- mented and not a 3-segmented club as noted in the original description. The similar species B. sijthofft, described also from Java, has small granules on the elytral inter- vals. Distribution: Java. Bradymerus elongatus (Perty, 1831) (Figs. 27, 92) Boletophagus elongatus Perty, 1831 (not Bradymerus elongatus Gebien, 1913). Bradymerus javanus Fairmaire, 1897 syn. Studied type material: None. New material: Java, leg. pe HAAN & Kayser, 4 ex. MNHUB (Hist. Coll. no. 47988). — E Java, Tengger Mt., 4000ft., leg. H. FRUHSTORFER, 1 ex. MNHUB. - W Java, Pengalengan, 4000ft., 1893, leg. H. FRUHSTORFER, 4 ex. MNHUB (det. GEBIEn). — Java, Soekaboemi, G. Malang, 1.1940, leg. OEko, 1 ex. HNHM. - Java, Gounod Gedeh, Ledru, 1898, 1 ex. SMNS (det. Kaszag). — Java, Preanger, Mt. Tangkoeban, Prahoe, 1800 m, XII.1935, 2 ex. NHMB-F (det. KuLZEr). — Java, Preanger, Mt. Tangkoeban, Prahoe, 1800 m, 1.1937, 2 ex. BMNH. - W Java, Gunung Gede-Pangrango NP, 10km NW Sukabumi, 1200m, 2.11.2001, leg. R. GERSTMEIER, 3 ex. CRGT. - Java, Mt. Kawi, ex coll. R. OBERTHÜR, 3 ex. HNHM. Diagnostic characters: Dorsal view see Fig.27, dorsal side ferrugineous without metallic shine, body length 8.0-9.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin somewhat sinuated, pronotal disc with rough punctation and with medial impression, between punctures without granules. Elytral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct keels, keels with granules, alternate intervals 2, 4, 6 with lower longitu- 16 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 dinal granules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 92. Distribution: Java. Bradymerus eschscholtzi Gebien, 1921 (Figs. 28, 93) Studied type material: Philippines, Luzon, Manila, leg. EscHscHoLTz, 1 d syntype MNHUB (labelled by GEBIEN as type, Hist. Coll. no. 46463), designated herewith as lecto- type. ae material: Philippines, Luzon, Camarines Sur, Mabatobato, Pili, 16.V.1931, 13 NHMB-F (det. Kutzer). Diagnostic characters: Dorsal view see Fig. 28, dorsal side blackish, with dis- tinct metallic shine, body length 9.5mm. Genae not broader than eyes, frons with- out distinct supraorbital keels but with deep supraorbital furrows. Last 5 anten- nomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and without medial impression, between punctures without granules. Elytral intervals 3 (posteri- orly), 5, 6, 7, 8 with keels, keels without granules. Tibiae in both sexes externally without distinct keels; in males all tibiae internally with a longitudinal furrow, in this furrow with a row of short, yellow setae. Aedeagus see Fig. 93. Distribution: Philippines (Luzon). Bradymerus ferruginipes Fairmaire, 1896 (Figs. 30, 95) Studied type material: None. New material: Philippines, Luzon, Camarines Sur, 12km N Ocampo, 800m, 25.-27.V.1996, leg. L. Boi, 4 ex. SMNS. - Philippines, Luzon, Mt. Banahao, leg. BoETT- CHER, 1 ex. ZSM (det. GEBIEN). — Philippines, Mindanao, Tangkulan, Bukidnon, leg. C. F. Baker, 1 ex. NHMB-F (det. GEBIEn). — Philippines, Mindanao, Davao, leg. C. F. BAKER, 1 ex. BMNH. - Philippines, Island Basilan near Mindanao, leg. BOETTCHER, 1 ex. MNHUB. - Philippines, Mindanao, Mt. Bango, leg. BOETTCHER, 1 ex. ZSM (det. Kaszas). — Philippines, Mindoro, 1 ex. BMNH (det. Bratr). — Borneo, Sabah, Ranau, 1500m, 1.VIII.1985, leg. K. Axıyama, 2 ex. HNHM. - Central Laos, Khammouan Prov., Ban Khoun Ngeun, 200m, 19.-31.V.2001, leg. L. DEMBICKY, 1 ex. SMNS. — Thailand, Chiang Dao, 350 m, 9.-14.V.1991, leg. D. KrAr & V. KuBAn, 11 ex. NHMB, 4 ex. SMNS. — NW Thailand, Mae Hong Song Prov., Ban Si Lang, 1200 m, 1.-8.V.1992, leg. J. HorAk, 1 ex. ZSM.- NW Thailand, Soppong, Pai, 1800 m, 1.-8.V.1993, leg. L. DEMBICKY & P. PACHOLATKO, 1 ex. ZSM. - NW Thailand, Nan Prov., Pha Knab, 11.-15.V1.1993, leg. L. DEMBICKY & P. PACHOLATKO, 1 ex. ZSM. - W Thailand, Khlong Lan NP, 2.-5.VI1.1997, leg. J. KALAB, 1 ex. ZSM. Diagnostic characters: Dorsal view see Fig. 30, dorsal side ferrugineous with- out metallic shine, body length 7.0-7.5 mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with dense punctation and with feeble impression, between punctures with fine granules. Elytral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct keels, keels with granules, alternate intervals 2, 4, 6 with low small granules. Tibiae in both sexes externally with indistinct keels. Aedeagus see Fig. 95. Remarks: GEBIEN (1913, 1925) treated this species only in the corresponding species-keys, but he gave no redescription, so probably he could not study type ma- terial. The new material from Luzon coincides with the specimen from Mindanao, identified by GEBIEN as Bradymerus ferruginipes, and even the zoogeographically remarkable newly collected specimens from Borneo, Thailand and Laos show no SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 17 distinct differences in all diagnostic characters. The species is quite similar to 2. propinquus, also from Mindanao. Distribution: Philippines (Luzon, Mindanao, Mindoro, Basilan); Borneo, Thailand, Laos (new records). Bradymerus fukiensis (Kaszab, 1954) n. comb. (Figs. 29, 96) Planibates fukiensis Kaszab, 1954. Studied type material: China, Fujian (labelled as Fukien), Kuatun, 2300 m, 1938, leg. J. KLAPPERICH, 2 dd paratypes of Planibates fukiensis HNHM. New material: Laos, Houaphan Province, Mt. Phu Phan (= Pan), 28.IV—6.V.2002, leg. H. Yosutromi, 18 CKAO. - Laos, Houaphan Province, Mt. Phu Phan, 1500-2000 m, 26.1V.-11.V.2001, leg. D. Hauck, 1 d SMNS. - Laos, Kiang Khonang, 1915, leg. R. V. DE SAL- vAZA, 1 ex. BMNH. - Vietnam (labelled as Tonkin), Mts. Mauson, 2000-3000ft., IV.-V. (year not labelled), leg. H. FRUHSTORFER, 222 MNHUB, 1d HNHM. - Vietnam (labelled as Tonkin), Chapa, 1915, leg. R. V. DE Satvaza, 1 ex. BMNH. Diagnostic characters: Dorsal view see Fig. 29, dorsal side blackish without metallic shine, body length 7.5-8.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 4 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with dis- tinct crenulation, pronotal disc with rough and confluent punctation and with me- dial impression, between punctures with granules. Elytral interval 5 with a row of el- evated granules or even granules confluent forming a keel, other intervals with a row of distinct granules, parly confluent on the intervals 3, 7. Tibiae externally without distinct keels; in males anterior tibia internally in the distal third with a distinct tooth. Aedeagus see Fig. 96. Remarks: Bradymerus fukiensis belongs to the small group of species with a modified anterior tibia in males and is therefore placed by Kaszaps in Planibates Kaszab, 1939. However, this character is not generic, all other characters coincide with Bradymerus. The type species of Planibates, Planibates papuanus Kaszab, 1939 from New Guinea, will be reexamined when treating the species of Bradymerus from New Guinea and adjacent islands. Distribution: China (Fujian, type locality); Laos, Vietnam (new records). Bradymerus grandis Fairmaire, 1893 Studied type material: None. New material: None. Diagnostic characters: The species is said to be 10.0mm long, the lateral margin of the pronotum not crenulated, elytra with rows of impressions (“fos- settes”) and all intervals with keels (FAIRMAIRE 1893). Distribution: Java. Bradymerus granulipennis Fairmaire, 1893 (Figs. 34, 100) Studied type material: None. New material: Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1560 m, 1.-5.1X.1988, leg. A. SMETANA, 2 ex. MHNG. - Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1560 m, 3.-13.VII1.1988, leg. A. SMETANA, 1 ex. MHNG. - Borneo, Sabah, Mt. Kinabalu NP, Head- quarters, Liwagu River, 1500 m, 18.V.1987, leg. A. SMETANA, 5 ex. MHNG, 3 ex. SMNS. - Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1500-1600m, 11.-15.X1.1996, leg. W. SCHAWALLER, 2 ex. SMNS. - Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1550m, 18 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 18.-25.V.2005, leg. R. Grimm, 31 ex. CRGT, 4 ex. SMNS. - Borneo, Sabah, Crocker Range, Gunung Emas, 1700m, 21.III.-20.IV.1996, leg. J. KapLEc, 31 ex. ZSM. — Borneo, Sabah, Crocker Range, Gunung Emas, 6.-18.V1.1996, leg. J. Kopapa, 2 ex. SMNS. - Borneo, Sabah, Keningau, 10.-20.X.1988, leg. M. Iron, 2 ex. CKAO. - Borneo, Sabah, Crocker Range, Gu- nung Emas, 15.-27.1V.1993, leg. M. Srrpa & I. Jeni, 21 ex. CSBC, 3 ex. SMNS. Diagnostic characters: Dorsal view see Fig. 34, dorsal side dark ferrugineous without metallic shine, body length 6.3-7.0mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation but somewhat sinuated, pronotal disc with rough punctation and with medial impres- sion, between punctures with granules. Internal elytral intervals with separated granules, external intervals with longitudinal, partly confluent granules. Tibiae in both sexes externally with distinct keels. Aedeagus see Fig. 100. Distribution: Borneo. Bradymerus impressicollis Gebien, 1913 (Figs. 37, 103) Studied type material: Philippines, Luzon, Benguet, Baguio, 4500-6000ft., leg. H. M. Curran, | d syntype NHMB-F (labelled by GEBIEN as type), designated herewith as lecto- type. New material: Philippines, Luzon, Benguet, Baguio, 1 ex. HNHM.- Philippines, N Lu- zon, Heightspian, leg. BOETTCHER, 1 ex. HNHM. - Philippines, N Luzon, Mountain Prov., Bontoc Region, NW Barlig, 2000 m, 9.TV.2000, leg. L. DEMBICKY, 2 ex. SMNS, 3 ex. HNHM, 2 ex. MNHUB, 2 ex. CKAO, 1 ex. CRGT. - Philippines, N Luzon, Kalinga-Apayao/Abra Province Boundery, Cordillera Central, 1600m, 26.-28.111.2000, leg. L. DEMBICKY, 6 ex. SMNS, 3 ex. CRGT, 1 ex. HNHM. - Philippines, Mindanao, 30 km NW Maramag, Bagongsi- lang, 1700 m, 13.-17.V.1996, leg. L. Borm, 17 ex. SMNS. — Philippines, Mindanao, Mt. Apo, Ilomavis, 1400 m, 18.-19.V.1996, leg. L. Borm, 1 ex. SMNS. — Philippines, Mindanao, 30 km W Maramag, 1600 m, 28.-30.X1I1.1990, leg. L. Boi, 2 ex. NHMB. - Philippines, Mindanao, N Sambuanga, Mt. Malindang, III.1995, local collector, 7 ex. HNHM. Diagnostic characters: Dorsal view see Fig. 37, dorsal side ferrugineous with- out metallic shine, body length 7.0-7.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, prono- tal disc with confluent punctation and with distinct medial impression, between punctures without granules. All elytral intervals with similar keels, keels without granules. Tibiae in both sexes externally without distinct keels; in males posterior tibia internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 103. Distribution: Philippines (Luzon, Mindanao). Bradymerus incostatus Gebien, 1914 (Figs. 38, 105) Studied type material: Simalur, Sinabang, V.1913, leg. E. Jacogson, 3 holotype NHMB-F (holotype by monotypy, labelled by GEBIEN as type). New material: NW Krakatau, IX.1920, 1 ex. HNHM. - Sumatra, Bengkhulu, 20 km S$ Muko Muko, 16.VIII.1991, leg. D. ERBER, 1 ex. SMNS. — Sumatra, Manna, 1901, leg. M. KNApPERT, 1 ex. BMNH. - Java, Jakarta (labelled as Batavia), III.1920, 1 ex. BMNH. - W Malaysıa, Johor, N Mersing, Kampung, Kaya Papan, 21.11.1994, leg. R. Grimm & A. RACHINSKY, 2 ex. CRGT.- W Malaysia, Nenasi, 18.VIII.1991, leg. U. BREMER, 1 ex. ZSM. Diagnostic characters: Dorsal view see Fig. 38, dorsal side ferrugineous with- out metallic shine, body length 6.5-7.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 19 corners of pronotum protruding, lateral margin without distinct crenulation, prono- tal disc with punctation and without medial impression, between punctures without granules. Elytral intervals slightly convex, without keels and without granules. Tib- iae in both sexes externally without distinct keels; in males posterior tibia internally near the tip with a field of distinct long, yellow setae. Aedeagus see Fig. 105. Distribution: Sunda Islands (Sumatra, Simalur, Nias, Krakatau, Java); W Malaysia (new record). Bradymerus interstitialis Schautuss, 1887 (Figs. 40, 107) Bradymerus alternatus Schaufuss, 1887 syn. Studied type material: S Celebes (Sulawesi), 1 d syntype of B. interstitialis MNHUB, designated herewith as lectotype of B. interstitialis. — S Celebes, 1 syntype of B. interstitialis NHMB-F (sex not examined), designated herewith as paralectotype of B. interstitialis. - S Celebes, 4 syntypes of B. alternatus MNHUB (sex not examined), designated herewith as lectotype and 3 paralectotypes of B. alternatus. — S Celebes, Bantimurang, 1882, leg. Ripe, 1 ex. MNHUB, might be a syntype of B. interstitialis (JAEGER in litt.). New material: S Sulawesi, Puncak Palopo, 2.1.2000, leg. K. Anno, 2 ex. CKAO, 2 ex. SMNS. Diagnostic characters: Dorsal view see Fig. 40, dorsal side ferrugineous with- out metallic shine, body length 8.0 mm. Genae broader than eyes, frons without dis- tinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and with medial impression, between punctures laterally with granules. Elytral intervals 3, 5, 7 with a fine keel, intervals 2, 4, 6 either with slightly lower and partly interrupted keels or with similar keels as on intervals 3, 5, 7, keels with granules. Tibiae in both sexes externally without distinct keels. Aedea- gus see Fig. 107. Distribution: Sumatra, Sulawesi, Sumbawa, Flores. Bradymerus iris Kulzer, 1951 (Figs. 43, 106) Studied type material: Philippines, Luzon, St. Thomas, leg. O. SCHÜTZE, d holotype NHMB-E New material: None. Diagnostic characters: Dorsal view see Fig. 43, dorsal side blackish, with dis- tinct metallic shine, body length 11.5 mm. Genae not broader than eyes, frons with- out distinct supraorbital keels but with deep supraorbital furrows. Last 5 anten- nomeres forming a club. Anterior corners of pronotum not protruding, lateral mar- gin without distinct crenulation, pronotal disc with separate punctation and without medial impression, between punctures without granules. Elytral intervals convex, without keels and without granules. Tibiae externally without distinct keels; in males posterior tibia internally near the tip with a field of distinct long, yellow setae. Aedeagus see Fig. 106. Distribution: Philippines (Luzon). Bradymerus kabakovi Kaszab, 1980 (Figs. 39, 108) Studied type material: China, Yunnan, Tamenglung, 1 paratype HNHM (sex not examined). New material: N Thailand, Mae Hong Son Prov., Ban Huai Po, 1800m, 20 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 30.1V.-14.V.1991, leg. J. FARKAC, 2 ex. SMNS. - N Thailand, Mae Hong Son Prov., Ban Si Lang, 1200 m, 23.-31.V.1991 and 1.-8.V.1992, leg. J. HorAx, 21 ex. ZSM, 2 ex. MHNL. - N Thailand, Chiang Mai Prov., NE Mae Taeng, Pong Deud Hot Springs, 30.TV.2003, leg. R. Grim, 11 ex. CRGT. - E Thailand, Chanthaburi Distr., Khao Soi Dao, 5.-13.V.1998, leg. J. HorAk, 1 ex. ZSM. — Thailand, Trat Prov., Ko Chang Island, 0-200 m, 1.-5.VII.2004, leg. R. & H. Fouquz, 6 ex. CRFL, 1 ex. SMNS. - S Thailand, Satun Prov., Thale Ban, 200 m, 8.-13.1V.1997, leg. J. KoLıgA&, 2 ex. NHMB. - Laos, Boli Kham Xai, 8km NE Ban Nape, 600 m, 1.-18.V.2001, leg. L. DEMBICKY, 1 ex. SMNS. — Indochina (Laos), between Ventiane and Luang Prabang, 1919, leg. R. V. DE Satvaza, 1 ex. BMNH. - Myanmar (labelled as Bur- ma), Mishmi Hills, Mondon, leg. M. STEELE, 1 ex. BMNH. — N India, Gopaldhara, 1915, leg. H. Stevens, 1 ex. BMNH. - NE India, Darjeeling, Rang, 26.IV.1987, leg. B. BHAKTA, 2 ex. NHMB. - India, Uttar Pradesh, Dehra Dun, Phanduwala, 4.V.1931, leg. J. C. M. GARDNER, 1 ex. BMNH. - Nepal, Sankhua Sabha Distr., Tumlingtar, 300 m, 28.V.1997, leg. M. Hauser & W. SCHAWALLER, 6 ex. SMNS. — Nepal, Chitwan Distr., Chitwan NP, Sauraha, 150 m, 31.V.-4.V1.1997, leg. M. Hauser & W. SCHAWALLER, 10 ex. SMNS. - Nepal, Chitwan NP, Sauraha, 20.-25.V.1992, leg. I. JENTS, 9 ex. ZSM. Diagnostic characters: Dorsal view see Fig. 39, dorsal side dark ferrugineous without metallic shine, body length 5.5-6.5 mm. Genae broader than eyes, frons with distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior cor- ners of pronotum protruding, lateral margin without distinct crenulation but some- what sinuated, pronotal disc with rough punctation, with medial impression and with a few high tubercles, between punctures with granules. Elytral intervals with high longitudinal granules, posteriorly confluent and forming a keel. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 108. Distribution: China (Yunnan), Vietnam (type localities); Nepal, India (Uttar Pradesh, Darjeeling), Thailand, Myanmar, Laos (new records). Bradymerus laevicostatus Kulzer, 1951 (Fig. 47) Studied type material: Philippines, NW Panay, leg. C. FE. Baker, 2 holotype NHMB- F (in bad condition, both antennae and legs partly missing). New material: None. Diagnostic characters: Dorsal view see Fig. 47, dorsal side blackish, with dis- tinct metallic shine, body length 12.0 mm. Genae not broader than eyes, frons with- out distinct supraorbital keels. Last 5 antennomeres forming a club (according to the description). Anterior corners of pronotum distinctly protruding, lateral margin without distinct crenulation, pronotal disc with separate punctation and without medial impression, between punctures without granules. Elytral intervals convex, without keels and without granules. Tibiae externally without distinct keels. Aedea- gus unknown, only female available. Distribution: Philippines (Island Panay). Bradymerus mcgregori Gebien, 1921 (Figs. 54, 120) Studied type material: Philippines, Luzon, Benguet, Irisan River, leg. McGreGor, 1 3 syntype NHMB-F (labelled by GrBIEN as type), designated herewith as lectotype. New material: Philippines, Luzon, Balbatan, leg. BOETTCHER, 1 ex. BMNH, 1 ex. HNHM. - Philippines, N Luzon, Kalinga-Apayao/Abra Province Boundery, Cordillera Central, 1600 m, 26.-28.111.2000, leg. L. Dempicxy, 11 ex. SMNS. - Philippines, Island Samar, Calbayog, VIII.1981, leg. SCHRAMM, 1 ex. CRGT. Diagnostic characters: Dorsal view see Fig. 54, dorsal side ferrugineous with- out metallic shine, body length 5.5-7.6mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 21 corners of pronotum protruding, lateral margin without distinct crenulation, prono- tal disc with rough punctation and with medial impression, between punctures with- out granules. Elytral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct keels, keels without granules, alternate intervals 2, 4, 6 with a few fee- ble granules. Tibiae in both sexes externally without distinct keels; in males last fe- mur internally before the middle and posterior tibıa internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 120. Remarks: The original and correct spelling of the species name is mcgregori and not macgregort. Distribution: Philippines (Luzon, Samar). Bradymerus mindanaensis Gebien, 1925 (Figs. 56, 116) Studied type material: Philippines, Mindanao, Kolambugan, leg. C. F. Baker, d holo- type NHMB-F (holotype by monotypy, labelled by GEBIEN as type). New material: Philippines, Mindanao, W Misamis, Don Victoriano, 1700m, 1.-3.V.1996, leg. L. Boi, 2 ex. SMNS. Diagnostic characters: Dorsal view see Fig. 56, dorsal side ferrugineous with- out metallic shine, body length 4.5-5.5 mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and with a feeble medial impression, between punctures without granules. Elytral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct keels, keels with granules. Tibiae in both sexes externally without dis- tinct keels. Aedeagus see Fig. 116. Distribution: Philippines (Mindanao). Bradymerus nodicollis Gebien, 1925 (Figs. 57, 122) Studied type material: Java, 1 2 syntype NHMB-F (labelled by GEBIEN as type), des- ignated herewith as lectotype. New material: Andaman Islands, without further dates, 1 ex. HNHM. - Borneo, Sabah, Crocker Range NP, NW Keningau, 900-1200 m, 16.-20.XI.1996, leg. W. SCHAWALLER, 1d SMNS. - Sumatra, Dolok Merangir, 28.VIII.1979, leg. D. ERBER, 1 ex. SMNS (det. Kaszas). — Java, Gounod, Kassi, 1 ex. HNHM. - S Sulawesi, Rante Pao, 9.-10.V1.1984, leg. G. ROUGE- MONT, 1 ex. HNHM. - S Sulawesi, Puncak Palopo, 2.1.2000, leg. K. ANDo, 1 ex. CKAO. Diagnostic characters: Dorsal view see Fig. 57, dorsal side ferrugineous with- out metallic shine, body length 5.5-7.5 mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation but somewhat sinuated, pronotal disc with rough punctation, with medial impression and with a pair of high tubercles, between punctures without granules. Alternate elytral inter- vals 3, 5, 7 with somewhat higher and complete keels, intervals 2, 4, 6 with somewhat lower and sometimes interrupted keels, keels with fine granules. Tibiae in both sex- es externally without distinct keels. Aedeagus see Fig. 122. Distribution: Andaman Islands, Java, Borneo, Sumatra, Sulawesi. Bradymerus pertyi Gebien, 1921 (Figs. 59, 124) Bradymerus elongatus Gebien, 1913 [secondary homonym of B. elongatus (Perty, 1831)]. Studied type material: Philippines, Luzon, Benguet, Panai, leg. MCGREGOR, 1 d syn- 22 STUTTGARTER BEITRAGE ZUR NATURKUNDE Ser. A, Nr. 694 type of B. elongatus Gebien, 1913 NHMB-F (labelled by GEBIEN as type), designated here- with as lectotype. New material: Philippines, Luzon, Benguet, Baguio, leg. C. F. Baker, 1 ex. HNHM (det. Kaszas). - Philippines, N Luzon, Heightspian, leg. BOETTCHER, 1 ex. HNHM. - Philip- pines, Luzon, Mt. Polis, leg. BOETTCHER, 1 ex. BMNH. - Philippines, Luzon, Baguio, Benguet, leg. C. F. Baker, 2 ex. BMNH. - Philippines, Luzon, Mountain Prov., Mt. Data, 5000ft., 19.11.1931, leg. F. C. HADDEN, 1 ex. HNHM (alternicostis det. KULZER). — Philip- pines, N Luzon, Mountain Prov., Botoc Region, NW Barlig, 2000 m, 9.IV.2000, leg. L. Dem- BICKY, 3 ex. SMNS. — Philippines, N Luzon, Zambales Mts., High Peak, westside, 850 m, 18.111.2000, leg. L. Drmpicxy, 3 ex. SMNS. - Philippines, Mindanao, Baracatan, 27.-29.V1.1977, leg. M. Sato, 4 ex. CKAO, 1 ex. SMNS. Diagnostic characters: Dorsal view see Fig. 59, dorsal side dark ferrugineous without metallic shine, body length 7.5-9.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation but somewhat sinuated, pronotal disc with confluent punctation and with medial impres- sion, between punctures without granules. Alternate elytral intervals 3 (only anteri- orly), 5, 7 with distinct keels, intervals 2, 4, 6 with separated distinct granules, keels with granules. Tibiae in both sexes externally without distinct keels; in males posteri- or tibia internally near the tip with a row of short, yellow setae. Aedeagus see Fig. 124. Distribution: Philippines (Luzon, Mindanao). Bradymerus planicollis Kulzer, 1951 (Fig. 60) Studied type material: Philippines, Luzon, Montalban, leg. O. SCHÜTZE, 2 holotype NHMB-E. New material: None. Diagnostic characters: Dorsal view see Fig. 60, dorsal side blackish without metallic shine, body length 9.0 mm. Genae not broader than eyes, frons without dis- tinct supraorbital keels. Last 5 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and without medial impression, between punctures with- out granules. Elytral intervals 3 (posteriorly), 5, 6 (anteriorly), 7 with keels, keels without granules. Tibiae externally without distinct keels. Aedeagus unknown, only female available. Distribution: Philippines (Luzon). Bradymerus propinquus Gebien, 1925 (Figs. 61, 125) Studied type material: Philippines, Mindanao, Tangkulan (labelled as Tangcolan), Bukidnon, leg. C. F. BAKER, d holotype NHMB-F (holotype by monotypy, labelled by GE- BIEN as type, in bad condition). New material: None. Diagnostic characters: Dorsal view see Fig. 61, dorsal side ferrugineous with- out metallic shine, body length 7.0mm. Genae broader than eyes, frons without dis- tinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with dense punctation and with feeble impression, between punctures with fine granules. All elytral intervals with distinct keels, keels without granules. Tibiae externally without keels. Aedeagus see Fig. 125. Distribution: Philippines (Mindanao). SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 23 Bradymerus seminitidus Arrow, 1900 (Figs. 65, 129) Studied type material: None. New material: Christmas Island, Flying Fish Cave, IX.1908, leg. C. W. ANDREws, 1 ex. NHMB-F (labelled by GEBIEN as “plesiotype” = topotype). — Christmas Island, Flying Fish Cave, IX.1908, leg. C. W. ANDREws, 3 ex. MNHUB. Diagnostic characters: Dorsal view see Fig. 65, dorsal side ferrugineous with- out metallic shine, body length 7.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with rough punctation and with medial impression, between punctures without granules. Elytral intervals 1, 2 with traces, all other intervals with similar distinct keels, keels with granules. Tibiae in both sexes externally without distinct keels; in males posterior tibia internally near the tip with a field of distinct long, yellow setae. Aedeagus see Fig. 129. Distribution: Christmas Island (380 km S Java). Bradymerus serricollis (Walker, 1858) (Figs. 66, 132) Studied type material: None. New material: Ceylon (Sri Lanka), leg. NIETNER, 4 ex. MNHUB (Hist. Coll. no. 47990). — Ceylon (Sri Lanka), Peradenyia, leg. FRIEDERICHs, 10 ex. MNHUB, 2 ex. SMNS. - Sri Lan- ka, Sinharaja, 5.X1I.1979, leg. V. MAHLER, 1 ex. HNHM. - Sri Lanka, 1 ex. NHMB-F. - Sri Lanka, Kandy, Udawattekele Sanctuary, 600 m, 24.XII.2000, leg. R. SCHUH, 2 ex. SMNS. - Sri Lanka, Ratnapura Distr., 2km S Hayes, 29.-30.X1.1995, leg. S. BECvAR & V. KosTAL, 5 ex. CSBC. - “Ind. or.”, 1 ex. HNHM. - China, Yunnan, 1 ex. HNHM. Diagnostic characters: Dorsal view see Fig. 66, dorsal side ferrugineous with- out metallic shine, body length 5.5-7.0 mm. Genae broader than eyes, frons with dis- tinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and without medial impression, between punctures with granules. Elytral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct keels, keels with granules. Tibiae in both sexes externally with distinct keels. Aedea- gus see Fig. 132. Remarks: In external morphology, this species is quite similar to Bradymerus crenulicollis from the Sunda Islands, but can be separated mostly by the different shape of the aedeagus (Figs. 91, 132), and by somewhat broader scale-like setae on the elytral intervals. See also B. thailandicus n. sp. Distribution: Sri Lanka (type locality); China (Yunnan) (new record). Bradymerus sijthoffi Gebien, 1925 (Figs. 67, 130) Bradymerus tibialis Kulzer, 1951 n.syn. Studied type material: Java, 1 d syntype of B. sijthofft MNHUB (labelled by GEBIEN as cotype, Hist. Coll. no. 46460), designated herewith as lectotype of B. sijthoffi. — Java, Pre- anger, leg. P. F. SyrHorr, 1 2 syntype of B. sjthoffi NHMB-F (labelled by GEBIEN as type), designated herewith as paralectotype of B. sijthoffi. - Java, Preanger, leg. P. F. SyrHorr, 1 syn- type of B. sijthoffi HNHM (labelled by Kaszas as Calabosca sijthoffi), designated herewith as paralectotype of B. sijthoffi. - The holotype of Bradymerus tibialis is not present in NHMB- F, and also no species-label in the corresponding drawer is present. New material: Borneo, Sabah, Kundasang, 1450 m, 14.-19.V.2005, leg. R. Grimm, 19 ex. CRGT, 4 ex. SMNS. - Borneo, Sabah, Kundasang, 27.1.2002, leg. R. ROBER, 1 ex. CRRH. - Borneo, Sabah, Crocker Range, route Keningau to Papar, V.1999, leg. M. SNIZEK, 1 ex. CMLS. 24 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 — Borneo, Sabah, Mt. Trus Madi, 1000 m, 16.1V.1996, leg. N. Kanıe, 1 ex. CKAO. - Borneo, Sabah, Mt. Trus Madi, 1200-1500 m, V.1995, leg. D. BoucHarp, 1 ex. CSBC. - Borneo, Sarawak, Kapit Distr., Rumah Ugab, Sut River, 3.-9.1II.1994, leg. J. HORAR, 1 ex. ZSM.- Bor- neo, Sarawak, Kapit Distr., Sebong, Baleh River, 9.-21.11I.1994, leg. J. HorAxk, 1 ex. CSBC, 1 ex. ZSM. - Borneo, Kalimantan, Busang/Rekut confluence, VIII.2001, leg. BRENDELL & MENDEL, 2 ex. BMNH. - S Sumatra, Lampung Prov., Bukit Barisan Selatan NP, 5 km SW Li- wa, 600 m, 7.-17.11.2000, leg. J. BEzpExk, 1 ex. SMNS. — Sumatra, Jambi, Gunung 'Tujhuh, Ker- inci NP, 1700-1900 m, 8.111.1991, leg. L. BocAx & M. BocAkovA, 1 ex. SMNS. — Sumatra, Gunung Tujuh, 5 km E Kersik Dua, 1900 m, 3.-5.V.2001, leg. L. Borm, 1 2 SMNS. - Sumatra, Brastagi, 900 m, 20.VII.1980, leg. E. Hetss, 1 ex. HNHM (labelled by Kaszag as Calabosca sp.). Diagnostic characters: Dorsal view see Fig. 67, dorsal side ferrugineous with- out metallic shine, body length 6.5-8.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 4 antennomeres forming a club. Anterior corners of pronotum slightly protruding, lateral margin without distinct crenulation but sinuated, pronotal disc with dense punctation and with medial impression, be- tween punctures without granules. All elytral intervals convex, interval 7 somewhat higher and with a row of fine pore-bearing granules. Tibiae in both sexes externally without distinct keels; in males anterior tibia internally in the distal third with a dis- tinct tooth. Aedeagus see Fig. 130. Synonymy: The sexually dimorphic male tibia in Bradymerus sijthoffi was not mentioned in the original description by GEBIEN (1925). This is just the character, which Kurzer (1951) considered as unique when describing B. tzbialis, also from Ja- va. All other characters given in the description of B. tibialis fully coincide with those in B. sijthoffi. Although the type of B. tibialis was not available for compari- son, I consider B. tibialis Kulzer, 1951 as a junior synonym of B. sijthoffi Gebien, 1925. Remarks: The antenna has a 4-segmented and not a 3-segmented club as noted in the original description. See also Bradymerus drescheri. Distribution: Java (type localities of B. sijthoffi and B. tibialis); Borneo, Suma- tra (new records). Bradymerus spretus Gebien, 1925 (Figs. 68, 131) Studied type material: Java, Buitenzorg, leg. A. PreyEr, 1d syntype NHMB-F (in bad condition, legs missing, labelled by GEBIEN as cotype), designated herewith as lectotype. — Same data, 1 2 syntype MNHUB, 1 syntype (only elytron) MNHUB, both designated herewith as paralectotypes. New material: S Sumatra, SW coast Ranau Lake, 1.-4.V1.2001, leg. L. Boım, 13 ex. NHMB, 4 ex. SMNS. — S Sumatra, Lampung, XI.1999, 2 ex. CKAO. - Borneo, Sabah, Keningau, 10.-20.X.1988, leg. M. Iron, 1 ex. CKAO, 1 ex. SMNS. — NW Thailand, Mae Hong Song, Ban Si Lang, 1000 m, 1.-7.V.1992, leg. S. Bıry, 1 ex. SMNS.- NW Thailand, Mae Hong Song, Ban Huai Po, 1800 m, 30.IV.-14.V.1991, leg. J. FarKac, 1 ex. NHMB. Diagnostic characters: Dorsal view see Fig. 68, dorsal side ferrugineous with- out metallic shine, body length 6.5-7.5 mm. Genae broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and with a feeble medial impression, between punctures without granules. All elytral intervals with similar fine keels, keels with small gran- ules. Tibiae in both sexes externally with fine keels. Aedeagus see Fig. 131. Distribution: Java; Sumatra, Borneo, Thailand (new records). SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 25 es a ere a ei ee a en i i Co Figs. 2-7. Dorsal views of Bradymerus species. — 2. B. acuticostis, 3 lectotype NHMB-E 3. B. acutangulus, 3 lectotype NHMB-F. 4. B. aeneus, 2 paratype HNHM. 5. B. aeratus, 2 holo- type NHMB-E 6. B. alternicostis, & holotype NHMB-E 7. B. andamanus, % lectotype NHMB-F. - Scale line: 5 mm. 26 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Figs. 8-13. Dorsal views of Bradymerus species. - 8. B. andoi n.sp., d holotype SMNS. 9. B. aratus, 3 holotype of syn. interruptus NHMB-F. 10. B. atronitens, 8 holotype NHMB-E. 11. B. asper, 2 holotype NHMB-F 12. B. aulacopterus, 3 lectotype ZFMK. 13. B. bifurcatus, 3 paratype HNHM. - Scale line: 5 mm. 27: SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS Figs. 14-19. Dorsal views of Bradymerus species. — 14. B. bocakorum n.sp., 3 holotype SMNS. 15. B. caeruleipennis, 2 holotype NHMB-F. 16. B. clathratus, 8 non-type SMNS. 17. B. crassicollis, 2 holotype NHMB-F. 18. B. crenulicollis, 8 non-type SMNS. 19. B. cari- natus, 6 non-type SMNS. - Scale line: 5 mm. 28 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 m 24 Figs. 20-26. Dorsal views of Bradymerus species. — 20. B. celebensis, 8 lectotype MNHUB. 21. B. difficilis, 8 holotype NHMB-F. 22. B. crockerensis n.sp., 6 holotype SMNS. 23. B. cu- cullatus, 2 non-type HNHM. 24. B. cyaneipennis, 2 non-type NHMB-F. 25. B. emasicus n.sp., d holotype SMNS. 26. B. drescheri, 3 lectotype NHMB-E - Scale line: 5 mm. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 29 Fe a it ake a ae Figs. 27-31. Dorsal views of Bradymerus species. — 27. B. elongatus, d non-type NHMB-F. 28. B. eschscholtzi, 3 non-type NHMB-F. 29. B. fukiensis, 3 non-type CKAO. 30. B. ferrug- inipes, 6 non-type SMNS. 31. B. fouquei n.sp., ¢ holotype CREL. - Scale line: 5 mm. 30 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Figs. 32-37. Dorsal views of Bradymerus species. — 32. B. gebieni n.sp., d holotype SMNS. 33. B. gerstmeieri n.sp., d holotype CRGT. 34. B. granulipennis, 3 non-type SMNS. 35. B. grimmi n.sp., d holotype CRGT. 36. B. hauseri n.sp., ¢ holotype SMNS. 37. B. impressicol- lis, 8 lectotype NHMB-F. - Scale line: 5 mm. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS al a i 13 f | I | | im Figs. 38-43. Dorsal views of Bradymerus species. — 38. B. incostatus, 3 holotype NHMB-F. 39. B. kabakovi, 3 non-type SMNS. 40. B. interstitialis, 3 lectotype MNHUB. 41. B. kanchenjungicus n. sp., d paratype SMNS. 42. B. kaszabin.sp., d holotype SMNS. 43. B. iris, 3 holotype NHMB-E - Scale line: 5 mm. Ser. A, Nr. 694 STUTTGARTER BEITRÄGE ZUR NATURKUNDE 32 a To 2 are os adi ok oak GA N = = to AZ A Be | on = 8. to 2 aed © Pie) “o . B. kulzeri n.sp., 48. B. masumotoi n.sp., & holotype MNST. 44-48. Dorsal views of Bradymerus species. — 44. B. kodadai n. sp 3 holotype SMNS. 46 a = 2 ae Ri Er EO a= 3) =e RS SS “8 88 Din & +S 5mm. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 33 Figs. 49-54. Dorsal views of Bradymerus species. — 49. B. laoticus n. sp., d holotype HNHM. 50. B. lombokicus n.sp., 3 holotype SMNS. 51. B. malayicus n.sp., 8 holotype SMNS. 52. B. majeri n.sp., d holotype HNHM. 53. B. maramagicus n.sp., d holotype SMNS. 54. B. mc- gregori, 3 lectotype NHMB-E - Scale line: 5 mm. 34 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Figs. 55-60. Dorsal views of Bradymerus species. — 55. B. michihikoi n.sp., 3 holotype SMNS. 56. B. mindanaensis, 8 holotype NHMB-E. 57. B. nodicollis, 2 lectotype NHMB-F. 58. B. merklin.sp., 2 holotype HNHM. 59. B. pertyi, 8 lectotype of syn. elongatus NHMB- F. 60. B. planicollis, 2 holotype NHMB-E - Scale line: 5 mm. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 35 Figs. 61-66. Dorsal views of Bradymerus species. - 61. B. propinquus, 3 holotype NHMB-F. 62. B. pseudomalayicus n.sp., 6 holotype SMNS. 63. B. reibnitzi n.sp., ¢ holotype SMNS. 64. B. riedeli n.sp., d holotype SMNS. 65. B. seminitidus, 3 non-type NHMB-F. 66. B. serri- collis, 3 non-type NHMB-E - Scale line: 5 mm. 36 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 73 Figs. 67-73. Dorsal views of Bradymerus species. — 67. B. sijthoffi, 3 lectotype MNHUB. 68. B. spretus, 3 lectotype NHMB-F. 69. B. sprecherae n.sp., d holotype SMNS. 70. B. suma- tranus n.sp., d holotype SMNS. 71. B. thailandicus n.sp., d holotype SMNS. 72. B. sum- bawaicus n.sp., ¢ holotype SMNS. 73. B. violaceus, 2 non-type SMNS. - Scale line: 5mm. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 85 Figs. 74-85. Aedeagi of Bradymerus species. — 74. B. acutangulus, 3 lectotype NHMB-F. 75. B. acuticostis, 3 lectotype NHMB-E. 76. B. alternicostis, 3 holotype NHMB-E 77. B. an- damanus, 8 non-type SMNS. 78. B. andoin.sp., ¢ holotype SMNS. 79. B. aratus, 3 holotype of syn. interruptus NHMB-E. 80. B. asper, 6 non-type BMNH. 81. B. bifurcatus, 3 paratype HNHM. 82. B. aulacopterus, 3 lectotype ZFMK. 83. B. atronitens, 8 holotype NHMB-F. 84. B. bocakorum n.sp., 3 holotype SMNS. 85. B. carinatus, d non-type SMNS. - Scale line: 1mm. il 37 38 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 92 9394 95 96 97 Figs. 86-97. Aedeagi of Bradymerus species. — 86. B. celebensis, 3 lectotype MNHUB. 87. B. clathratus, 8 non-type SMNS. 88. B. crockerensis n.sp., d holotype SMNS. 89. B. drescheri, 3 lectotype NHMB-F. 90. B. difficilis, 8 holotype NHMB-F. 91. B. crenulicollis, 8 non-type SMNS. 92. B. elongatus, 3 non-type CRGT. 93. B. eschscholtzi, 8 non-type NHMB-F. 94. B. emasicus n.sp., 6 holotype SMNS. 95. B. ferruginipes, 8 non-type SMNS. 96. B. fukiensis, 3 non-type CKAO. 97. B. fouquei n.sp., d holotype CREL. - Scale line: 1 mm. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 39 98 104 100 99 103 101 102 105 106 107 108 109 110 Figs. 98-110. Aedeagi of Bradymerus species. — 98. B. gebienin.sp., ¢ holotype SMNS. 99. B. gerstmeieri n.sp., d holotype CRGT. 100. B. granulipennis, 3 non-type SMNS. 101. B. grim- min.sp., 6 holotype CRGT. 102. B. hauseri n.sp., ¢ paratype CRGT. 103. B. impressicollis, 3 lectotype NHMB-E. 104. B. kanchenjungicus n.sp., ¢ holotype NHMB. 105. B. incostatus, 3 non-type CRGT. 106. B. iris, d holotype NHMB-F. 107. B. interstitialis, 3 lectotype MNHUB. 108. B. kabakovi, 3 non-type SMNS. 109. B. kaszabi n.sp., 5 holotype SMNS. 110. B. kinabalicus n.sp., & holotype SMNS. - Scale line: 1 mm. 40 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 an AAR 117 118 119 120 121 122 Figs. 111-122. Aedeagi of Bradymerus species. — 111. B. kodadai n.sp., d holotype SMNS. 112. B. kulzerin.sp., ¢ holotype SMNS. 113. B. lombokicus n. sp., d holotype SMNS. 114. B. malayicus n.sp., d holotype SMNS. 115. B. laoticus n.sp., ¢ holotype HNHM. 116. B. min- danaensis, 3 holotype NHMB-E. 117. B. majeri n.sp., d holotype HNHM. 118. B. maram- agicus n.sp., d holotype SMNS. 119. B. masumotoi n.sp., d holotype MNST. 120. B. mcgre- gori, 3d lectotype NHMB-F. 121. B. merkli n.sp., d paratype SMNS. 122. B. nodicollis, 3 non-type SMNS. - Scale line: 1 mm. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 41 i 129 123 it 128 130 131 132 133 134 135 136 Figs. 123-136. Aedeagi of Bradymerus species. — 123. B. michihikoi n.sp., 3 holotype SMNS. 124. B. pertyi, 3 lectotype of syn. elongatus NHMB-F. 125. B. propinquus, 3 holotype NHMB-F. 126. B. pseudomalayicus n.sp., d holotype SMNS. 127. B. reibnitzi n. sp., d holo- type SMNS. 128. B. riedeli n.sp., d holotype SMNS. 129. B. seminitidus, d non-type NHMB-F. 130. B. sijthoffi, 8 lectotype MNHUB. 131. B. spretus, 3 lectotype NHMB-F. 132. B. serricollis, ¢ non-type HNHM. 133. B. sumatranus n.sp., & holotype SMNS. 134. B. sprecherae n.sp., d holotype SMNS. 135. B. sumbawaicus n.sp., 6 holotype SMNS. 136. B. thailandicus n.sp., ¢ holotype SMNS. - Scale line: 1 mm. 42 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Bradymerus violaceus Pascoe, 1883 (Fig. 73) Studied type material: None. New material: Philippines, Luzon, Benguet, 1 2 NHMB-F (det. Kurzer). — Philip- pines, Sibuyan, 12 HNHM (det. Kaszas). — Philippines, Sibuyan, Romblon, 1 ex. CKAO, 1 ex. SMNS. - Philippines, N Luzon, Kalinga-Apayao/Abra Province Boundery, Cordillera Central, 1600 m, 26.-28.111.2000, leg. L. DEmBIcKY, 1 2 SMNS. Diagnostic characters: Dorsal view see Fig. 73, dorsal side blackish, with dis- tinct metallic shine, body length 8.5-9.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation, prono- tal disc with rough punctation and without medial impression, between punctures without granules. Internal elytral intervals with smaller, external intervals with big- ger separated granules. Tibiae externally without distinct keels. Aedeagus unknown, only females available. Distribution: Philippines (Luzon, Sibuyan). 4 Descriptions of new Oriental species of Bradymerus Bradymerus andoi n.sp. (Figs. 8, 78) Holotype (4): Central Sulawesi, Palu Palolo, Lindu NP, 25.-27.VIIl.1990, leg. A. RIEDEL, SMNS. Paratypes: S Sulawesi, Puncak Palopo, 1.-2.1.2000, leg. K. ANDo, 4 ex. CKAO. - S Su- lawesi, Kalosi, Alla Enrekang, 29.XT1.1999, leg. K. Anno, 1 ex. SMNS. — S Sulawesi, Puncak Palopo, 20.1.2000, leg. G. BeccE, 1 ex. CKAO. - Sulawesi, Selatan, To’Rea, 1.2000, local col- lector, 1 ex. CKAO. - Central Sulawesi, Poso, 5-10km SW ‘Tambarana, 400m, 11.-16.1V.1999, leg. S. BECVAR & P. ZABRANSKY, 1 ex. CSBC. - S Sulawesi, 35 km NW Palopo, 1400 m, 18.IV.1999, leg. S. BECVAR & P. ZABRANSKY, 2 ex. CSBC. Etymology: Named in honor of Dr. KtyosH1 ANDO (Osaka), one of the collectors of the type series, for long-term and fruitful cooperation. Description: Dorsal view see Fig. 8, dorsal side blackish without metallic shine, body length 7.8-9.0 mm. Genae not broader than eyes, frons without distinct supra- orbital keels, with distinct supraorbital furrows. Last 5 antennomeres forming a club. Anterior corners of pronotum distinctly protruding, lateral margin with feeble crenulation, pronotal disc with rough but separate confluent punctation, pronotal disc without medial impression, only laterally between punctures without granules. Internal elytral intervals 1-3 with a row of separate longitudinal granules, external intervals with confluent longitudinal granules forming keels. Tibiae in both sexes externally with distinct keels. Aedeagus see Fig. 78. Diagnosis: To be recognized by the shape of the pronotum with distinctly pro- truding anterior corners, by the dorsal structure of the elytra and by the shape of the aedeagus. Bradymerus acutangulus from Java and B. sumatranus n. sp. from Sumatra have a similar shape of the pronotum, but their body size is smaller (6.0-6.5 mm), and the aedeagi are different (Figs. 74, 78, 133). Bradymerus bocakorum n.sp. (Figs. 14, 84) Holotype (6): Philippines, Mindanao, 30 km E Malaybalay, Busdi, 1000 m, 5.-9.V.1996, leg. L. Bo-m, SMNS. Etymology: Named in honor of Dr. Mırapa BocAkovA and Dr. Lapistav BocAK SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 43 (Olomouc) for deposing their collectings of Coleoptera from different travels through Asia in SMNS. Description: Dorsal view see Fig.14, dorsal side blackish without metallic shine, body length 6.5mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 5 antennomeres forming a club. Anterior corners of pronotum only slightly protruding, lateral margin without distinct crenulation, pronotal disc with distinct and sometimes confluent punctation and without medial impression, between punctures without granules. Alternate ely- tral interval 3 only posteriorly with a low keel, intervals 5, 7 with complete and high keels, keels with granules, internal intervals flat and with some indistinct granules. Tibiae externally without distinct keels. Aedeagus see Fig. 84. Diagnosis: Bradymerus bocakorum n. sp. belongs to the small group of species with a 5-segmented antennal club around B. acutangulus from Java, B. bifurcatus from Vietnam, B. maramagicus n.sp. also from Mindanao and B. sumatranus n. sp. from Sumatra. These taxa can be separated by a different shape of the pronotum and by a different structure of the elytra (Figs. 3, 13, 14, 53, 70) as well as by a different shape of the aedeagus (Figs. 74, 81, 84, 118, 133). Bradymerus crockerensis n.sp. (Figs. 22, 88) Holotype (d): Borneo, Sabah, Crocker Range NP, NW Keningau, 900-1200m, 16.-20.X1.1996, leg. W. SCHAWALLER, SMNS. Etymology: Named after the Crocker Mt. Range, where the holotype was collected. Description: Dorsal view see Fig.22, dorsal side blackish without metallic shine, body length 9.0mm. Genae not broader than eyes, frons without distinct supraorbital keels, with weak supraorbital furrows. Last 4 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation but slightly sinuated, pronotal disc with confluent punctation and with medial impression, between punctures without granules. Elytral intervals convex, al- ternate intervals 3, 5, 7 slightly higher but without distinct keels, intervals with a row of indistinct pore-bearing granules. Tibiae in both sexes externally without distinct keels; in males anterior tibia internally in the middle with a distinct tooth. Aedeagus see Fig. 88. Diagnosis: To be recognized by the sexually dimorphic anterior tibia with an internal tooth in the middle in males, by long antenna (reaches the posterior margin of the pronotum) with a 4-segmented antennal club and by the structure of the ely- tra without any distinct keels. Bradymerus drescheri and B. sijthoffi from the Sunda Islands are the hitherto known species also possessing a modified anterior male tib- ia, but the tooth is situated in the distal third, the elytra have the interval 7 nearly keel-like and the antenna is short (reaches only the middle of the pronotum). See al- so B. hauseri n. sp. from the Malayan Peninsula; in this species the tooth of the male anterior tibia is situated in the distal third as in B. drescheri, but the elytra possess distinct alternate keel-like intervals. The aedeagus is quite similar in this species- group (Figs. 88, 89, 102, 130). See also B. kinabalicus n. sp. from Borneo with a 3-seg- mented antennal club and the pronotum with distinct tubercles between the rough punctation, and B. grimmi n. sp. also from Borneo. 44 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Bradymerus emasicus n.sp. (Figs. 25, 94) Holotype (d): Borneo, Sabah, Crocker Range, Gunung Emas. 6.-18.V1.1996, leg. J. Ko- DADA, SMNS. Paratypes: Same data as holotype, 1 ex. SMNS. - Borneo, Sabah, km 53 road Kota Kina- balu to Tambunan, Gunung Emas, 1650m, 6.IV.2000, leg. L. Borm, 5 ex. NHMB, 2 ex. SMNS. - Borneo, Sabah, km 53 road Kota Kinabalu to Tambunan, E slope Gunung Emas, 700 m, 1.-5.IV.2000, leg. L. Borm, 1 ex. NHMB. - Borneo, Sabah, Crocker Range, Gunung Emas, 15.-27.1V.1993, leg. I. JEnı$ & M. Srrpa, 1 ex. ZSM. - Borneo, Sabah, Crocker Range, Gunung Emas, 500-1900 m, 6.-21.V.1995, leg. I. JENIS, 1 ex. ZSM. — Borneo, Sabah, Crocker Range, 16km SW Gunung Alab, 790-850 m, 4.-9.V.1996, leg. M. Srrpa & R. Hercovirs, Tex CSBC Etymology: Named after Gunung (= Mount) Emas, where the type series was collected. Description: Dorsal view see Fig. 25, dorsal side ferrugineous without metallic shine, body length 5.5-7.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin without distinct crenulation but sinuated, pronotal disc with confluent punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3, 5, 7 with keels, keels with granules, intervals 4, 6 with a row of longitudinal granules, mostly confluent and forming a lower keel. Tibiae in both sexes externally without distinct keels; in males posterior tibia internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 94, Diagnosis: To be recognized by the small body size, the shape and dorsal struc- ture of the pronotum and elytra, by the 6-segmented antennal club, by the modified posterior tibiae in males, and by the shape of the aedeagus. The combination of these characters is not comparable in any other Oriental species. Bradymerus fouquei n.sp. (Figs. 31, 97) Holotype (d): W Malaysia, Pahang, Gunung Jasar, Tanah Rata, 1500-1700m, 2.-4.V11.2001, leg. R. Fouquéz & H. Bartovd, CREL. Paratypes: Same data as holotype, 2 ex. CRFL. - W Malaysia, Pahang, Gunung Jasar, Tanah Rata, 1500-1700 m, 8.-17.VIl.2004, leg. R. & H. Fouqus, 2 ex. CRFL, 2 ex. SMNS. Etymology: Named in honor of REn£ Fouqui (Liberec), collector of the type series. Description: Dorsal view see Fig.31, dorsal side blackish without metallic shine, body length 7.5-8.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with weak supraorbital furrows. Last 4 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation but slightly sinuated, pronotal disc with confluent punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3 (widely interrupted in the middle), 5, 7 with keels, keels with granules, intervals 1, 2, 4, 6 with a row of granules. Tibiae in both sexes externally without distinct keels; in males anterior tibia internally in the basal third with a pair of distinct teeth. Aedea- gus see Fig. 97. Diagnosis: Bradymerus fonquei n. sp. is extremely similar to B. hauseri n. sp. al- so from W Malaysia, but can be recognized by a darker and less shining dorsal side, by a longer pronotum with rougher punctation (Figs. 31, 36) and particularly by the male anterior tibia internally with a pair of distinct teeth in the basal third, which is as yet unique within the genus (in B. hauseri n.sp. and other species only a single tooth in the distal third), and by the different shape of the aedeagus (Figs. 97, 102). SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 45 Bradymerus gebieni n.sp. (Figs. 32, 98) Holotype (d): W Sumatra, Bengkhulu Prov., near Curup, Bukit Kaba Mt., 1000-1500 m, 30.1.-2.11.2000, leg. J. BEZDER, SMNS. Paratypes: Same data as holotype, 6 ex. SMNS. — Sumatra, Brastagi, leg. MJOBERG, 3 ex. NHMB-F. — W Sumatra, Gunung Singgalang, 1800m, VII.1925, leg. E. Jacogson, 1 ex. MNHUB (elongatus Perty det. GEBIEN). Etymology: Named in memory of Hans GEBIEN (1874-1947), first monographer (1925) of the genus Bradymerus. Description: Dorsal view see Fig. 32, dorsal side ferrugineous without metallic shine, body length 5.5-6.8 mm. Genae slightly broader than eyes, frons with distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and with medial impression, between punctures with granules. All ely- tral intervals with longitudinal and confluent tubercles forming interrupted keels, al- ternate intervals 3, 5, 7 slightly more prominent. Tibiae in both sexes externally with distinct keels. Aedeagus see Fig. 98. Diagnosis: To be recognized by the shape and dorsal structure of the pronotum and elytra, by only slightly dilated genae, by a 6-segmented antennal club, and by the shape of the aedeagus. Extremely similar, also in the shape of the aedeagus, is B. aratus Fairmaire, 1896 from Borneo and Sumatra, but in this species the genae are distinctly dilated, the pronotum is more convex and the tubercles on the elytral in- tervals are much higher and not confluent. As long as no intermediate forms are known, I consider both as different species. See also B. gerstmeieri n. sp. from Suma- tra. Bradymerus gerstmeieri n.sp. (Figs. 33, 99) Holotype (4): Sumatra, Prov. Jambi, Kerinci-Seblat NP, Danau Tujuh, 2000-2250 m, 11.-14.VII.2001, leg. R. GERSTMEIER, CRGT. Paratypes: Same data as holotype, 1 ex. CRGT, 1 ex. SMNS. Etymology: Named in honor of Prof. Dr. ROLAND GERSTMEIER (Freising), collector of the type series. Description: Dorsal view see Fig. 33, dorsal side ferrugineous without metallic shine, body length 7.0-7.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin with feeble crenulation, pronotal disc with rough punctation and with medial impression, between punctures without granules. All elytral intervals with longitudinal and confluent tubercles forming interrupted keels, alternate intervals 3, 5, 7 slightly more prominent. Tibiae in both sexes externally with distinct keels. Aedeagus see Fig. 99. Diagnosis: Bradymerus gerstmeieri n. sp. can be recognized by the 6-segmented antennal club, by the undilated genae, by the shape and dorsal structure of the pronotum and elytra, and by the shape of the aedeagus. Quite similar in external morphology is B. gebieni n.sp., also from Sumatra, but apart from a smaller body size (5.5-6.8 mm in B. gebieni n.sp.), this species has slightly dilated genae, distinct supraorbital keels, a different shape and structure of the pronotum (Figs. 32, 33), and a different shape of the aedeagus (Figs. 98, 99). 46 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Bradymerus grimmi n.sp. (Figs. 35, 101) Holotype (4): Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1550 m, 22.-25.V.2005, leg. R. Grimm, CRGT. Paratypes: Borneo, Sabah, Crocker Range NP, Gunung Emas, 6.-18.V1.1996, leg. J. Ko- DADA, 1 2 SMNS. Etymology: Named in honor of Dr. ROLAND Grim (Tübingen), one of the collectors of the type series, for long-term and fruitful cooperation. Description: Dorsal view see Fig. 35, dorsal side blackish without metallic shine, body length 6.5-7.2 mm. Genae not broader than eyes, frons without distinct supra- orbital keels, with weak supraorbital furrows. Last 4 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with crenulation, pronotal disc with partly confluent punctation and with medial impression, between punctures without granules. All elytral intervals convex, intervals with a row of pore-bearing granules. Tibiae in both sexes externally without distinct keels; in males anterior tibia internally in the middle with a distinct tooth. Aedeagus see Fig. 101. Diagnosis: Bradymerus grimmi n. sp. is quite similar to B. crockerensis n. sp. al- so from Borneo, but apart from a smaller body size (9.0 mm in B. crockerensis n. sp.), the shape of the pronotum is different (Figs. 22, 35), the granules on the elytral in- tervals are more distinct and more densely situated, and the aedeagus is different (Figs. 88, 101). Bradymerus hauseri n.sp. (Figs. 36, 102) Holotype (4): W Malaysia, Cameron Highlands, Tanah Rata, 2.VIII.1999, leg. M. Hauser, SMNS. Paratypes: Same data as holotype, 2 ex. SMNS. — W Malaysia, Pahang, Cameron High- lands, 30 km E Ipoh, Tanah Rata, 1500 m, 20.I1.-3.1I1.1998, leg. P. CecHovskyY, 1 ex. SMNS. - W Malaysia, Cameron Highlands, Tanah Rata, 1500m, 14.-18.111.1998, leg. P. CecHovsky, 8 ex. ZSM.- W Malaysia, W Perak, 25 km NE Ipoh, Banjaran Titi Wangsa Mts., Mt. Korbu, 1200 m, 6.-12.V.2001, leg. P. CecHovskY, 2 ex. SMNS. — W Malaysia, Pahang, Cameron High- lands, Parit Falls, 27.111.1995, leg. O. MERKL, 4 ex. HNHM. - W Malaysia, Pahang, Cameron Highlands, Tanah Rata, foothills of Gunung Beremban, 29.1I1.1995, leg. O. MERKL & G. CsorBa, 1 ex. HNHM. - W Malaysia, Cameron Highlands, Tanah Rata, 1500m, 13.-20.1V.1999, leg. A. BALLERIO, 1 ex. CRSW. — W Malaysia, Kelantan, 30 km NE Tanah Ra- ta, 800 m, 17.-19.1V.1999, leg. A. BALLERIO, 1 ex. CRSW, 1 ex. SMNS. - W Malaysia, Pahang, Cameron Highlands, Tanah Rata, 9.11.1994, leg. R. Grimm & A. RACHINSKY, 8 ex. CRGT. - W Malaysia, Tanah Rata, 28.V.1975 and 22.V1.1975, leg. Y. Kıyoyama, 2 ex. CKAO. - W Malaysia, Pahang, Gunung Jasar, Tanah Rata, 1500-1700 m, 29.V1.-4.V11.2001, leg. R. Fouqut & H. BARLOVA, 43 ex. CRFL, 6 ex. SMNS. — W Malaysia, Pahang, Gunung Jasar, Tanah Rata, 1500-1700 m, 8.-17.VII.2004, leg. R. & H. Fouqué, 28 ex. CRFL, 6 ex. BMNH. — W Malaysia, Pahang, Gunung Brinchang, Brinchang, 1600-2000m, 3.VII.2001, leg. R. Fouqur & H. BARLOVA, 18 ex. CRFL. - W Malaysia, Pahang, Gunung Brinchang, Brin- chang, 1600-2000 m, 11. and 14.VII.2004, leg. R. & H. Fouquz, 4 ex. CRFL. — W Malaysia, Pahang, Cameron Highlands, Tanah Rata, Gunung Jasar, 1400-1500 m, 20.-25.1.1995, leg. S. BECvAR, 6 ex. CSBC. — W Malaysia, Pahang, Cameron Highlands, Tanah Rata, Gunung Jasar, 1400-1500 m, 19.-25.V1.1995, leg. S. BECvAR, 10 ex. CSBC. — W Malaysia, Pahang, Cameron Highlands, Tanah Rata, Gunung Jasar, 1400-1500 m, 12.-15.11.1998, leg. S. BECVAR, 10 ex. CSBC. - W Malaysia, Pahang, Cameron Highlands, Brinchang, Gunung Beremban, 1600 m, 18.-19.1.1995, leg. S. BECvVAR, 4 ex. CSBC. — W Malaysia, Perak, Taiping, Bukit Larut (Maxwell Hill), 14.1V.1996, leg. S. BECVAR, 1 ex. CSBC. — W Malaysia, Perak, 1 ex. BMNH. - W Malaysia (labelled as Malay. Peninsula), Kedah Peak, 3000ft., 8.III.1928, leg. H. M. PENDLEBURY, 1 ex. BMNH. Etymology: Named in honor of Dr. Martin Hauser (Urbana/lllinois), one of the col- lectors of the type series, for fruitful cooperation and deposing several of his tenebrionids in Stuttgart. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 47 Description: Dorsal view see Fig.36, dorsal side brownish without metallic shine, body length 6.8-8.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with weak supraorbital furrows. Last 4 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin without distinct crenulation but slightly sinuated, pronotal disc with confluent punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3 (widely interrupted in the middle), 5, 7 with keels, keels with granules, intervals 1, 2, 4,6 with a row of granules. Tibiae in both sexes externally without distinct keels; in males anterior tibia internally in the distal third with a distinct tooth. Aedeagus see Fig. 102. Diagnosis: To be recognized by the sexually dimorphic anterior tibia in males with an internal tooth in the distal third, by a 4-segmented antennal club, by the structure of the elytra with distinct keels on the alternate intervals. Bradymerus drescheri and B. sijthoffi from the Sunda Islands are the only hitherto known species possessing also a similarily modified anterior male tibia, but the structure of the ely- tral intervals (Figs.26, 36, 67), and also the shape of the aedeagus is different (Figs. 89, 102, 130). A sexually dimorphic male tibia is also present in B. fukiensis from Indochina and Fujian, in B. crockerensis n.sp. and B. kinabalicus n.sp. both from Borneo, but these species have also a different dorsal structure on pronotum and elytra, a different shape of the aedeagus, and B. kinabalicus n.sp. has a 3-seg- mented antennal club. See also under B. laoticus n.sp. from Laos and B. fouquei n. sp. from Malaysia. Bradymerus kanchenjungicus n.sp. (Figs. 41, 104) Holotype (3): E Nepal, Kanchenjunga Himal, Chiruwa, 1260 m, 30.VI.-1.VII.2000, leg. J. FARKAG, D. KrAL & J. SCHNEIDER, NHMB. Paratypes: Same data as holotype, 2 ex. NHMB, 1 ex. SMNS. — E Nepal, Taplejung Distr., lower Gunsa Khola to Lungthung, 1650-1870 m, 18.V.1988, leg. J. MarTENs & W. SCHAWALLER, 1 2 SMNS. — E Nepal, Arun Valley, Lamobagar, 1400 m, 8.-14.V1.1983, leg. C. HoLzscHuH, 2 ex. MHNL. Etymology: Named after Mt. Kanchenjunga in northeastern Nepal, where the type series was collected nearby. Description: Dorsal view see Fig.41, dorsal side blackish without metallic shine, body length 7.0-7.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 5 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation but sinuated, pronotal disc with confluent punctation and without me- dial impression, between punctures without granules. All elytral intervals convex, without keels and granules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 104. Diagnosis: To be recognized by the elytral intervals without keels and granules and by the broad pronotum without protruding anterior corners and with confluent punctation, and without modified posterior tibiae in males. Bradymerus incostatus from the Sunda Islands and W Malaysia possesses similar unmodified elytral inter- vals without keels and without granules, but the pronotum has distinctly protruding anterior corners and a finer punctation, and a sexual dimorphic posterior tibia inter- nally near the tip with a field of distinct long, yellow setae in males. The aedeagi of both taxa have similar acute joint parameres (Figs. 104, 105). See also B. malayicus n. sp. 48 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Bradymerus kaszabi n.sp. (Figs. 42, 109) Holotype (6): S Sulawesi, 20km NE Sabbang, 400m, 5.-7.VI1.2001, leg. L. Boi, MNS. Paratypes: Same data as holotype, 5 ex. SMNS, 2 ex. HNHM. - S Sulawesi, 8 km W Ma- masa, 950m, 18.—21.VII.1999, leg. L. Borm, 1 ex. SMNS. - S Sulawesi, 15km W Palopo, 11.-19. VIII.1990, leg. A. RıeDEL, 1 ex. SMNS. — Sulawesi, Kotamogabu, Matalibaru, Torosik, Gunung Tongara, 700 m, 10.XII.1999, leg. A. Rıeper, 1 ex. SMNS. - Central Sulawesi, Toar- co Jaya, Rante Pao, 2.V1.1984, leg. M. Tao, 1 ex. NSMT. - S Sulawesi, Puncak Palopo, Luwu Palopo, To’Rea, 18.-23.1.2000, leg. G. BEccE, 4 ex. CKAO. - S Sulawesi, Puncak Palopo, Luwu Palopo, km 27, 20.1.2000, leg. G. BEcce, 4 ex. CKAO. - Sulawesi, Selatan, To’Rea, 1.2000, local collector, 4 ex. CKAO. —S Sulawesi, Puncak Palopo, 2.1.2000, leg. K. ANDO, 2 ex. CKAO. - S Sulawesi, Puncak Palopo, 30.XI1.1999, leg. Y. UrsuNomrya, 2 ex. CKAO. - S Sulawesi, Puncak Palopo, 18.-19.1.2000, leg. B. Gata, 1 ex. SMNS. Etymology: Named in memory of Dr. ZOLTAN Kaszag (1915-1986), former director of the Hungarian Natural History Museum, member of the Hungarian Academy of Sciences, coleopterologist of world-wide recognition and outstanding specialist of the Tenebrionidae. Description: Dorsal view see Fig.42, dorsal side dark ferrugineous without metallic shine, body length 6.5-8.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with distinct crenulation, pronotal disc with rough and confluent punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3, 5, 7 with complete keels, intervals 2, 4, 6 with identical keels but only in the anterior part of the elytra, keels with gran- ules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 109. Diagnosis: To be recognized by the 6-segmented antennal club, by the shape of the pronotum with protruding anterior corners with a crenulated lateral margin and with rough and confluent dorsal punctation, by the structure of the elytra with iden- tical keels on all intervals, and by the shape of the aedeagus. The combination of these characters is not comparable in any other Oriental species. See also Bradymerus lombokicus n. sp. Bradymerus kinabalicus n.sp. (Figs. 45, 110) Holotype (4): Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1500-1600 m, 11.-15.X1.1996, leg. W. SCHAWALLER, SMNS. Paratypes: Same data as holotype, 1 ex. SMNS. — Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1550 m, 22.-25.V.2005, leg. R. Grimm, 3 ex. CRGT. — Borneo, Sabah, Mt. Kin- abalu NP, Poring Hot Springs, 600 m, 20.VIII.1998, leg. D. BarrscH & C. HÄUSER, 1 ex. SMNS. - Borneo, Sabah, Mt. Kinabalu, 28.V.1999, leg. Z. SMRR, 1 ex. ZSM. — Borneo, Sabah, Mt. Kinabalu, Lumu Lumu, 5500ft., 12.1V.1929, leg. H. M. PENDLEBURY, 1 ex. BMNH. - Borneo, Sabah, Crocker Range, Gunung Emas, 1600m, 13.V.2005, leg. R. Grimm, 3 ex. CRGT. - Borneo, Sabah, Crocker Range NP, Gunung Emas, 6.-18.V1.1996, leg. J. Kopapa, 1 ex. SMNS. — Borneo, Sabah, Crocker Range, Gunung Alab, 1700 m, 23.-29.V.1998, leg. J. Kopapa & F. Crampor, 1 ex. SMNS. - Borneo, Sabah, Crocker Range, Gunung Emas, 15.-27.1V.1993, leg. I. Jeni & M. Srrpa, 6 ex. ZSM. — Borneo, Sabah, km 53 on road Kota Kinabalu to Tambunan, Gunung Emas, 1650 m, 22.III.-6.IV.2000, leg. L. Bom, 3 ex. NHMB. — Borneo, Sabah, km 53 on road Kota Kinabalu to Tambunan, E slope Gunung Emas, 700 m, 1.-5.IV.2000, leg. L. Bor, 1 ex. SMNS. — Borneo, Sabah, Gunung Emas, 23.-29.V.1998, leg. P. HravA&, 1 ex. HNHM. - Borneo, Sabah, Mt. Kinabalu, W slope, 1900 m, 5.II1.1969, leg. H. LÖFFLER, 1 ex. HNHM. - Borneo, Sabah, Kundasang, 27.1.2002, leg. R. ROBER, 1 ex. CRRH. — Borneo, Sabah, Tibow, 45km NE Sapulut, 600-900 m, 7.-15.IV.2000, leg. L. Boi, 1 ex. CSBC. - Borneo, Sabah, Gunung Emas 42 km E Kota Kinabalu, 21.III.-20.IV.1996, leg. Lin- DA, 4 ex. CSBC. — Borneo, Sabah, Crocker Range, Gunung Alab, 1650-1800 m, 30.1V.-27.V.1996, leg. M. Srrpa & R. Hercovirs, 4 ex. CSBC. Etymology: Named after Mt. Kinabalu, where the type series was collected nearby. SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 49 Description: Dorsal view see Fig.45, dorsal side blackish without metallic shine, body length 7.5-8.0 mm. Genae elevated but not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 3 anten- nomeres forming a club. Anterior corners of pronotum slightly protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and with me- dial impression, between punctures with distinct tubercles. All elytral intervals with a row of distinct granules, granules on alternate intervals 3, 5, 7 denser and higher. Tibiae in both sexes externally without distinct keels; in males anterior tibia inter- nally in the distal third with a distinct tooth. Aedeagus see Fig. 110. Diagnosis: To be recognized by the sexually dimorphic anterior tibia in males with an internal tooth in the distal third, by long antennae with a 3-segmented an- tennal club, by the structure of the pronotum with distinct tubercles between the rough punctation, by the elytral structure with rows of granules and by the shape of the aedeagus. Other species with a modified anterior tibia in males (for example Bradymerus drescheri, B. sijthoffi, B. crockerensis n.sp. and B. hauseri n.sp.) have a 4-segmented antennal club, a pronotum without granules or tubercles between the finer punctation, a different structure of the elytra, and a different aedeagus. Bradymerus kodadai n.sp. (Figs. 44, 111) Holotype (8): Borneo, Sabah, Crocker Range NP, Gunung Emas, 6.-18.V1.1996, leg. J. Kopapa, SMNS. Paratypes: Same data as holotype, 1 ex. SMNS. — Borneo, Sabah, Crocker Range, Gu- nung Emas, 15.-27.1V.1993, leg. I. JENS & M. Srrpa, 10 ex. ZSM. — Borneo, Sabah, Gunung Emas, 23.-29.V.1998, leg. P. HravA&, 1 ex. HNHM. - Borneo, Sabah, Crocker Range, Gu- nung Alab, 1700m, 23.-29.V.1998, leg. J. Kopapa & F. Crampor, 2 ex. SMNS. - Borneo, Sabah, Mt. Kinabalu NP, Headquarters, 1550 m, 22.-25.V.2005, leg. R. Grimm, 1 ex. CRGT. - Borneo, Sabah, Mt. Kinabalu, Lumu Lumu, 5500ft., 15.1V.1929, leg. H. M. PENDLEBURY, 1 ex. BMNH. - Borneo, Sarawak, 19.-20.111.1995, leg. M. Iron, 1 ex. CKAO. Etymology: Named in honor of Dr. JAN Kopapa (Bratislava), one of the collectors of the type series, for deposing several of his tenebrionids in Stuttgart. Description: Dorsal view see Fig.44, dorsal side dark ferrugineous without metallic shine, body length 6.5—7.5 mm. Genae not broader than eyes, frons with- out distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior cor- ners of pronotum slightly protruding, lateral margin slightly sinuated, pronotal disc with confluent punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3, 5, 7 with higher and confluent lon- gitudinal granules forming keels, intervals 2, 4, 6 with somewhat lower and more separate granules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 111. Diagnosis: Bradymerus kodadai n.sp. shares with B. elongatus (Perty, 1831) from Java the undilated genae and the general structure of the elytra with the higher alternate intervals, but can be separated by a different shape of the pronotum (Figs. 27, 44; broader and flat in B. elongatus, narrower and more convex in B. ko- dadai n.sp.), and by a slightly different shape of the aedeagus (Figs. 92, 111). Quite similar in external morphology and also in the shape of the aedeagus is B. sum- bawaicus n.sp. from Sumbawa, Bali and Sulawesi. The shape and structure of the pronotum and the structure of the elytral intervals in these three taxa are similar but not identical, also when considering a certain variability. Thus, I decided to treat these disjunct insular populations (B. elongatus from Java, B. kodadai n.sp. from 50 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Borneo and B. sumbawaicus n.sp. from Sumbawa, Balı and Sulawesi) as different taxa, as long as no intermediate forms are known. Bradymerus kulzeri n.sp. (Figs. 46, 112) Holotype (d): Central Sulawesi, 20km SE Tambarana, Camp Mauro, 11.-16.V11.1999, leg. L. Bo-m, SMNS. Paratypes: Same data as holotype, 6 ex. SMNS, 2 ex. HNHM. - Central Sulawesi, 38 km SE Pendolo, 1200 m, 10.-11.VII.2001, leg. L. Borm, 3 ex. SMNS, 2 ex. MNHUB. - S Sulawe- si, 25km E Mamasa (Kalama), 1100 m, 1.-3.V11.2001, leg. L. Bot, 1 ex. SMNS. - S Sulawesi, 8km W Mamasa, 950 m, 18.-21.V11.1999, leg. L. BoLm, 1 ex. SMNS. - Central Sulawesi, Toar- co Jaya, Rante Pao, 2.-5.V1.1984, leg. M. Tao, 8 ex. NSMT. — Sulawesi, Puncak Palopo, 22.X.-3.X1.1985, leg. M. Tao, 2 ex. NSMT. — Sulawesi, Puncak Dingin, 17.X1.1985, leg. M. Tao, 3 ex. NSMT. — S Sulawesi, Puncak Palopo, 2.1.2000, leg. K. ANDo, 1 ex. CKAO. - S Sulawesi, Puncak Palopo, Luwu Palopo, To’Rea, 18.1.2000, leg. G. BEccE, 1 ex. CKAO. - Sulawesi, Selatan, To’Rea, 1.2000, local collector, 1 ex. CKAO. - S Sulawesi, Tanah Toraja, Karum-Ganga, 27.XI1.1999, leg. M. Anno, 9 ex. CKAO, 3 ex. SMNS. - S Sulawesi, Rante Pao, 9.-10.V1.1984, leg. G. ROUGEMONT, 1 ex. HNHM. - Central Sulawesi, 15-25 km S Pen- dolo, Mayoa, 7.-10.IV.1999, leg. S. BECvAR & P. ZABRANSKY, 5 ex. CSBC. — Central Sulawesi, Poso, 5-10km SW Tambarana, 400 m, 11.-16.IV.1999, leg. S. BECVAR & P. ZABRANSKY, 2 ex. CSBC. - Central Sulawesi, W Coast of Lake Poso, Taipa, 10.-11.1V.1999, leg. S. BECvAR & P. ZABRANSKY, 11 ex. CSBC. — Central Sulawesi, 20-35km NW Palopo, 1000-1400 m, 4.-5.1V.1999, leg. S. BECVAR & P. ZABRANSKY, 1 ex. CSBC. — S Sulawesi, Puncak Palopo, V.1999, local collector, 3 ex. CSBC. — W Sulawesi, Bambapuang Pass, road Rante Pao to Rap- pang, 29.X.2005, leg. E. Heıss, 1 ex. CRGT. Etymology: Named in memory of Hans Kurzer (1889-1974), former curator of the Frey collection, for his substantial contribution upon the genus Bradymerus (1951) and oth- er papers about Oriental and Neotropical Tenebrionidae. Description: Dorsal view see Fig. 46, dorsal side dark ferrugineous without metallic shine, body length 7.3-8.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with rough and confluent punctation and without medial impression, between punctures with granules. All elytral intervals wrinkled by rough punctation, internal intervals flat and anteriorly with a row of indistinct granules, external intervals with indistinct keels. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 112. Diagnosis: To be recognized by undilated genae, by the rough punctation of the pronotum without protruding anterior corners and without crenulated lateral mar- gin, by the structure of the elytra with flat but densely wrinkled internal intervals, and by the shape of the aedeagus. In particular the flat but wrinkled elytral intervals are quite characteristic and do not occur in any other as yet known species. Bradymerus laoticus n.sp. (Figs. 49, 115) Holotype (8): Laos, Champassak Prov., Dong Hua Xao NBCA, 2km S Ban Nong Lu- ang, bank of Touay-Guai River, 800 m, 1.-5.1V.1998, leg. O. MERKL & G. Csorsa, HNHM. Paratypes: Same data as holotype, 10 ex. HNHM, 3 ex. SMNS. Etymology: Named after Laos, where the type series was collected. Description: Dorsal view see Fig.49, dorsal side blackish without metallic shine, body length 8.0-9.8 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 4 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 51 crenulation but slightly sinuated, pronotal disc with confluent punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3,5, 7 with identical keels, keels without granules, intervals 1, 2, 4, 6 flat and without distinct granules. Tibiae in both sexes externally without distinct keels; in males an- terior tibia internally in the distal third with a distinct tooth. Aedeagus see Fig. 115. Diagnosis: Bradymerus laoticus n.sp. shares with B. hauseri n.sp. from W Malaysia the armed anterior male tibia and the general body shape and size, but can be separated by a somewhat bigger body size in the average (6.8-8.5 mm in B. hauserin.sp.), by a dull dorsal surface (shining in B. hauseri n. sp.), by high and iden- tical keels on the elytra (compare Figs. 36, 49) and by a different shape of the aedea- gus (compare Figs. 102, 115). B. merkli n.sp. from Vietnam with similar body size possesses a similar dull surface, but besides other characters (aedeagus, Fig. 121) the elytral keels are present only on the intervals 5, 7 and the punctures of the elytral in- tervals are of longitudinal shape and deeply impressed. Bradymerus lombokicus n.sp. (Figs.50, 113) Holotype (4): Lombok, Sapit-Sembalun Bumbung, 900-1500 m, 14.-16.11.1994, leg. L. BoLm, SMNS. Paratypes: Same data as holotype, 2 ex. SMNS. Etymology: Named after the island Lombok, where the type series was collected. Description: Dorsal view see Fig. 50, dorsal side ferrugineous without metallic shine, body length 8.5-9.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin without distinct crenulation, pronotal disc with dis- tinct and sometimes confluent punctation and with medial impression, between punctures without granules. Alternate elytral intervals 3, 5, 7 with complete keels, intervals 2, 4, 6 with identical keels but only in the anterior part of the elytra, keels with granules. Tibiae in both sexes externally with or without distinct keels. Aedea- gus see Fig. 113. Diagnosis: Bradymerus lombokicus n. sp. shares with B. kaszabi n. sp. from Su- lawesi the undilated genae, the 6-segmented antennal club, the dorsal structure of the elytra, and also the shape of the aedeagus (Figs. 109, 113). Besides the bigger body size (body length 6.5-8.0 mm in B. kaszabi n.sp.), the shape and dorsal structure of the pronotum is completely different (Figs. 42, 50). Bradymerus majeri n.sp. (Figs. 52, 117) Holotype (¢): India, Andaman Islands, Island Cinque (no. 5), 27.XII.1978, leg. G. OsEL- La, HNHM. Paratypes: India, Andaman Islands, Island Havelock, around village no. 7, 22.1V.-14.V.1998, leg. K. & S. Majer, 3 ex. NHMB, 1 ex. SMNS. Etymology: Named in memory of KareL Majer (1949-2000), one of the collectors of the type series and well-known Czech Coleopterologist. Description: Dorsal view see Fig.52, dorsal side blackish without metallic shine, body length 7.5-9.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 5 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation, pronotal disc with confluent punctation and without medial impres- sion, between punctures without granules. Elytral intervals 3 (posteriorly), 4 (poste- 52 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 riorly), 5, 6, 7 with keel, keels with granules, internal intervals 1-4 on the elytral disc flat, with fine punctation, without granules but somewhat wrinkled. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 117. Diagnosis: To be recognized by the broad pronotum without protruding ante- rior corners, by the structure of the elytra, and by the shape of the aedeagus. Bradymerus pertyi Gebien, 1921 from the Philippines has a somewhat similar struc- ture of the elytra, but in this species the flat internal intervals bear some granules and only the alternate intervals are keel-like, the pronotum has a different shape and rougher punctation, and the aedeagus is different (Figs. 117, 124). Bradymerus malayicus n.sp. (Figs. 51, 114) Holotype (3): W Malaysia, Pahang, Cameron Highlands, 30km E Ipoh, Tanah Rata, 1500 m, 22.-26.1.1999, leg. P. CacHovsxy, SMNS. Paratypes: Thailand, Chumphon Prov., Pha To, 1.-20.1I1.1996, leg. K. Mayer, 1 ex. SMNS. — Thailand, Chumphon Prov., Pha To, 14.III.-14.IV.1996, leg. P. PRÜDER, 30 ex. CSBC, 8 ex. SMNS, 2 ex. HNHM. - India, Assam, U. Dihing, Lakhimpur, 28.V.1921, leg. C. F. C. BEESON, 1 ex. BMNH. Etymology: Named after the Malayan Peninsula, where the holotype was collected. Description: Dorsal view see Fig.51, dorsal side dark ferrugineous without metallic shine, body length 6.0-6.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation but sinuated, pronotal disc with fine and separate punctation and without medial impression, be- tween punctures without granules. Elytral intervals slightly convex, without keels and without granules. Tibiae in both sexes externally without distinct keels. Aedea- gus see Fig. 114. Diagnosis: To be recognized by the 5-segmented antennal club, by convex ely- tral intervals without keels and granules, by the flat and broad pronotum with fine and separate punctation and without protruding anterior corners, by unmodified posterior tibiae in males and by the shape of the aedeagus. A similar elytral structure has Bradymerus incostatus Gebien, 1914 from the Sunda Island and W Malaysia, and B. kanchenjungicus n. sp. from Nepal. B. incostatus, however, has a different shape of the pronotum with distinctly protruding anterior corners and modified posterior tibiae in males, and B. kanchenjungicus n.sp. has also a different shape of the prono- tum widest before the middle and with denser confluent punctation (in B. malayicus n.sp. pronotum widest in the middle and with fine and separate punctation). See al- so B. pseudomalayicus n. sp. from the same area. Bradymerus maramagicus n.sp. (Figs.53, 118) Holotype (6d): Philippines, Mindanao, 30km W Maramag, 1600 m, 28.-30.XII.1990, leg. L. BoLm, SMNS. Etymology: Named after the village Maramag, in whose vicinity the holotype was col- lected. Description: Dorsal view see Fig.53, dorsal side blackish without metallic shine, body length 7.2 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 5 antennomeres forming a club. Anterior corners of pronotum distinctly protruding, lateral margin without distinct crenulation, pronotal disc with distinct and sometimes confluent punctation and without medial impression, between punctures with indistinct granules. Elytral SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS => intervals with distinct granules, granules on intervals 5, 7 sometimes confluent and forming interrupted keels. Tibiae externally without distinct keels. Aedeagus see Fig. 118. Diagnosis: Bradymerus maramagicus n.sp. belongs to the small group of species with a 5-segmented antennal club around B. acutangulus from Java, B. bifur- catus from Vietnam, B. bocakorum n.sp. also from Mindanao and B. sumatranus n.sp. from Sumatra. These taxa can be separated by a different shape of the prono- tum and a different structure of the elytra (Figs. 3, 13, 14, 53, 70) and by a different shape of the aedeagus (Figs. 74, 81, 84, 118, 133). Bradymerus masumotoi n.sp. (Figs. 48, 119) Holotype (¢): Tatwan, Island Lan Hsu (= Lanyu, = Orchid Island), Tienchi, 400m, 2.V.2005, leg. K. Masumoto, J.-F. Tsar & W.-Z. CHEN, MNST. Paratypes: Same data as holotype, 12 SMNS. - Taiwan, Island Lan Hsu, 5.-8.V1.1986, leg. S. Osawa, 1? NHMC. Etymology: Named in honor of Dr. Kınıo Masumoto (Tokyo), one of the collectors of the type series, for long-term and fruitful cooperation. Description: Dorsal view see Fig. 48, dorsal side blackish with a feeble metallic shine, body length 9.0-10.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin with a feeble crenulation, pronotal disc with separate punctation and without impression, between punctures without granules. Elytral in- tervals slightly convex, alternate intervals 3, 5, 7 posteriorly with feeble keels, all in- tervals without granules. Tibiae in both sexes externally without distinct keels; in males posterior tibia internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 119. Diagnosis: Bradymerus masumotoi n. sp. is extremely similar to B. aulacopterus from Saleyer Island adjacent of Sulawesi, but can be separated by a somewhat nar- rower pronotum slightly excavated before the hind angles and by a feeble crenula- tion of the lateral margin (compare Figs. 12, 48), by a feeble metallic shine of the dor- sal surface (without metallic shine in B. aulacopterus) and by a somewhat different shape of the aedeagus (Figs. 82, 119). A further argument for the validity of both species might be the wide disjunct distribution (B. masumotoi n. sp. near Taiwan, B. aulacopterus near Sulawesi). Bradymerus merkli n.sp. (Figs. 58, 121) Holotype (¢): Vietnam, Lam Dong Prov., Da Lat, 17.X.1988, leg. S. MaHUNKA & T. VASARHELYI, HNHM. Paratypes: Vietnam (labelled as Annam), Da Lat (labelled as Dalat), 23.1I1.-9.1V.1924, leg. R. V. DE Satvaza, 2 ex. BMNH, 1 ex. SMNS. Etymology: Named in honor of Dr. Orrö Merkr (Budapest), who recognized this species already in 1991 as new, for fruitful cooperation and hospitality during my visits. Description: Dorsal view see Fig.58, dorsal side blackish without metallic shine, body length 7.5-8.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 4 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin without distinct crenulation but sinuated, pronotal disc with separate punctation and with distinct medial impression, between punc- tures without granules. Elytral intervals 5, 7 with keels, keels without granules, in- ternal intervals 1-4 on disc flat, longitudinal punctures of the rows deeply im- 54 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 pressed. Tibiae in both sexes externally without distinct keels; in males anterior tib- ia internally in the distal third with a distinct tooth. Aedeagus see Fig. 121. Diagnosis: To be recognized by the 4-segmented antennal club, by the shape and punctation of the pronotum, and by the structure of the elytral intervals. Bradymerus merkli n.sp. belongs to the species-group around B. crockerensis n. sp. from Borneo, B. fukiensis from Indochina and Fujian and B. hauseri n.sp. from W Malaysia with a 4-segmented antennal club and with sexually dimorphic anterior tibiae in males, but cannot be mixed up because of the unique dorsal structure of pronotum and elytra. See also under B. laoticus n.sp. and B. fouquei n. sp. Bradymerus michihikoi n.sp. (Figs. 55, 123) Holotype (¢): Central Sulawesi, 38km SE Pendolo, 1200 m, 10.-11.VII.2001, leg. L. BoLm, SMNS. Paratypes: Sulawesi, 31 km from Palopo, 5.V1.1982, leg. M. Tao, 2 2? NSMT. - Sulawe- si, Sampuraga, 4.X1.1985, leg. M. Tao, 12 NSMT. — Sulawesi, Selatan, Puncak Palopo, 2.1.2000, leg. N. OHBAYASHI, 2 ex. CKAO. - Sulawesi, Puncak Palopo, 2.1.2000, leg. M. An- bo, 1 ex. CKAO. Etymology: Named in honor of MicHIHIko ANDO (Osaka), son of Dr. KrıyosHı ANDo, one of the collectors of the type series. Description: Dorsal view see Fig.55, dorsal side blackish without metallic shine, body length 8.8-9.5 mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep and extraordinary wide supraorbital furrows. Last 5 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin with feeble crenulation, pronotal disc with rough and confluent punctation, pronotal base before scutellum with fine and separate punctation, pronotal disc without medial impression, between punctures without granules. Alternate elytral intervals 3, 5, 7 with higher, intervals 4, 6, 8 with somewhat lower keels, keels with- out granules. Tibiae in both sexes externally without distinct keels; in males posteri- or tibia internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 123. Diagnosis: To be recognized by the deep and extraordinary wide supraorbital furrows in both sexes, by the humped pronotum with rough and confluent puncta- tion on the disc and with fine and separate punctation at the base, by keeled elytral intervals and by the shape of the aedeagus. The combination of these characters does not occur in any other of the compared species. Bradymerus pseudomalayicus n.sp. (Figs. 62, 126) Holotype (8): W Malaysia, Perak, 25 km NE Ipoh, Banjaran Titi Wangsa Mts., Mt. Kor- bu, 1200 m, 1.-15.IV.2000, leg. P. CecHovsky, SMNS. Paratypes: Same data as holotype, 1 ex. SMNS. — W Malaysia, Perak, 25 km NE Ipoh, Banjaran Titi Wangsa Mts., Mt. Korbu, 1200 m, 6.-12.V.2001, leg. P. CecHovskyY, 1 ex. SMNS. — Borneo, Sarawak, Kapit Distr., Rumah Ugap, Sut River, 3.-9.11I.1994, leg. J. HorAx, 3 ex. CSBC, 1 ex. ZSM. - W Borneo (labelled as Borneo occ.), Gunung (labelled as Goenong) Am- par, 1897, leg. MuLoT, 2 ex. HNHM (labelled by Kaszas as Calabosca sp.). Etymology: Named because of the extreme similarity to Bradymerus malayicus n. sp. from the same area. Description: Dorsal view see Fig.62, dorsal side dark ferrugineous without metallic shine, body length 5.5-7.3 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 4 antennomeres forming a club. Anterior corners of pronotum not protruding, lateral margin without distinct crenulation but sinuated, SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 55 pronotal disc with separate punctation and with a feeble medial impression, between punctures without granules. Elytral intervals slightly convex, without keels and without granules. Tibiae in both sexes externally without distinct keels. Aedeagus see Fig. 126. Diagnosis: To be recognized by the 4-segmented antennal club, by convex ely- tral intervals without keels and granules, by the shape of the pronotum without pro- truding anterior corners, by unmodified posterior tibiae in males and by the shape of the aedeagus. Bradymerus psendomalayicus n.sp. is extremely similar to B. malayi- cus n.sp. from the same area, but has a 4-segmented antennal club (5-segmented in Bradymerus malayicus n.sp.), a distinctly longer pronotum with bigger and denser punctation (Figs.51, 62), a dull dorsal surface (shining in Bradymerus malayicus n.sp.), and a diseniery different shape of the aedeagus (Figs. 114, 126). Bradymerus reibnitzi n.sp. (Figs. 63, 127) Holotype (8): Central Java, Gunung Slamet, 5km N Baturaden, 1100 m, 12.-13.V.2001, leg. L. Bo-m, SMNS. Etymology: Named in honor of JOHANNES REIBNITZ (Stuttgart) for his professional efforts in preparing the photographs and plates. Description: Dorsal view see Fig. 63, dorsal side blackish with distinct metallic shine, body length 7.5mm. Genae not broader than eyes, frons without distinct supraorbital keels, with deep supraorbital furrows. Last 4 antennomeres forming a club. Anterior corners of pronotum protruding, lateral margin with feeble crenula- tion, pronotal disc with separate punctation and with feeble transverse mediobasal impression, between punctures without granules. Elytral intervals flat and without keels or granules, punctural rows in striae. Tibiae externally without distinct keels; in males anterior tibia internally in the distal third with distinct tooth and distinctly bent inwards, posterior tibia swollen behind the middle and internally before apex with distinct excavation. Aedeagus see Fig. 127. Diagnosis: Bradymerus reibnitzi n.sp. belongs to the group of species with a 4-segmented antennal club and with an armed anterior tibia in males, but it is the on- ly Oriental species of the genus not only with modified anterior but also with mod- ified posterior tibia in males. Additionally, this species can be recognized by the dis- tinct metallic dorsal surface, by the shape of the pronotum with protruding anterior corners, by the elytral intervals without granules and keels, and by the shape of the aedeagus. Remarks: The long and narrow shape of the aedeagus (Fig. 127) differs extraor- dinary from all other congeners. This might be a further hint that Bradymerus in the present scope is not monophyletic (see introduction). Bradymerus riedeli n.sp. (Figs. 64, 128) Holotype (d): S Sulawesi, Tanah Toraja, Pulu Pulu, Ponding, 13.-16.VIII.1990, leg. A. RIEDEL, SMNS. Paratypes: Same data as holotype, 2 ex. SMNS. Etymology: Named in honor of Dr. ALEXANDER RIEDEL (Karlsruhe), the collector of the type series, for fruitful cooperation and depositing several of his tenebrionids in Stuttgart. Description: Dorsal view see Fig.64, dorsal side dark ferrugineous without metallic shine, body length 7.8-8.0 mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 5 antennomeres forming a club. Anterior corners of 56 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 pronotum protruding, lateral margin without distinct crenulation but sinuated, widely separated and distinctly bent upwards, pronotal disc with rough punctation and with a feeble medial impression, between punctures without granules. Alternate elytral intervals 3, 5, 7 with interrupted, feeble keels, keels with granules, intervals 2, 4, 6 without granules, punctures of the rows deeply impressed. Tibiae in both sexes externally without distinct keels; in males posterior tibia internally in the middle with a row of short, yellow setae. Aedeagus see Fig. 128. Diagnosis: To be recognized by the shape of the pronotum with the lateral mar- gin round, widely separated and distinctly bent upwards, by undilated genae and frons without distinct supraorbital keels, by the structure of the elytra, and by the shape of the aedeagus. This is the only Oriental species of the genus with such a shape of the pronotum. A similar pronotum possesses Bradymerus buruensis Kulz- er, 1951 from the Moluccas, but this species has dilated genae, distinct supraorbital keels, a distinct surface of the pronotum with tubercles, distinct elytral intervals without keels; the aedeagus is unknown. Bradymerus sprecherae n.sp. (Figs. 69, 134) Holotype (4): Central Sulawesi, Palu Palolo, Lindu NP, 25.-27.VIII1.1990, leg. A. RIEDEL, SMNS. Etymology: Named in honor of Dr. Eva SPRECHER (Basel) for fruitful cooperation and hospitality during my visits. Description: Dorsal view see Fig. 69, dorsal side ferrugineous without metallic shine, body length 3.5mm. Genae about as broad as eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and with deep medial impression, between punctures with distinct tu- bercles. Elytral interval 3 in the middle with high keel, intervals 4-7 with distinct longitudinal granules partly confluent and forming incomplete keels, keels with granules. Tibiae externally with distinct keels. Aedeagus see Fig. 134. Diagnosis: To be recognized by the minute body size (smallest as yet known species of the genus), by the 6-segmented antennal club, by the shape and dorsal structure of pronotum and elytra, by the keeled tibiae, and by the shape of the aedeagus. It shares with Bradymerus asper from the Philippines the relatively small body size (4.5mm), the 6-segmented antennal club and the keeled tibiae, but both can be distinguished by the shape of the pronotum and the different structure of the elytra (Figs. 11, 69), and by the shape of the aedeagus (Figs. 80, 134). Bradymerus sumatranus n.sp. (Figs. 70, 133) Holotype (d): N Sumatra, Pematang-Siantar, 18km direction to Prapat, 1000m, IV.-IX.1996, leg. E. W. Dienr, SMNS. Etymology: Named after the island Sumatra, where the type was collected. Description: Dorsal view see Fig.70, dorsal side dark ferrugineous without metallic shine, body length 6.0 mm. Genae not broader than eyes, frons without dis- tinct supraorbital keels, with deep supraorbital furrows. Last 5 antennomeres form- ing a club. Anterior corners of pronotum distinctly protruding, lateral margin with- out distinct crenulation but sinuated, pronotal disc with dense punctation and with- out medial impression, between punctures laterally with a few granules. Elytral intervals 3 (only posteriorly), 4-8 with keels, keels with granules, internal intervals SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 57 1-3 on the disc with a row of granules. Tibiae in both sexes externally without dis- tinct keels. Aedeagus see Fig. 133. Diagnosis: To be recognized by the shape of the pronotum with distinctly pro- truding anterior corners, by the structure of the elytral intervals, and by the shape of the aedeagus. Bradymerus sumatranus n.sp. shares with B. acutangulus from Java the relatively small body size (6.0-6.5 mm), the 5-segmented antennal club, the deep supraorbital furrows, and the distinctly protruding anterior corners of the prono- tum (Figs.3, 70). Both can be mainly separated by the completely different dorsal structure of the elytra (Figs.3, 70). The shape of the aedeagus is similar in both species (Figs. 74, 133), but quite different from the other congeners. See also B. bo- cakorum n.sp. and B. maramagicus n.sp., both from Mindanao. Bradymerus sumbawaicus n.sp. (Figs. 72, 135) Holotype (8): W Sumbawa, Batudulang, 30km S Sumbawa Besar, 1000 m, 10.11.1994, leg. L. Bo-m, SMNS. Paratypes: Same data as holotype, 26 ex. SMNS, 2 ex. CKAO, 2 ex. CRGT, 2 ex. HNHM, 2 ex. MNHUB, 2 ex. NHMB. - Central Sulawesi, 20km SE Tambarana, Camp Mauro, 650 m, 11.-16.VII.1999, leg. L. Borm, 2 22 SMNS. — S Sulawesi, Bantimurang, 1882, leg. G. Risse, 1 ex. MNHUB. - S Sulawesi, Puncak Palopo, 2.1.2000, leg. K. Anno, 1 ex. CKAO. - S Sulawesi, Puncak Palopo, Salu-Bua, 18.-19.1.2000, leg. B. Gara, 1 ex. CKAO. - S Sulawesi, Puncak Palopo, 30.XII.1999, leg. Y. UrsuNomtya, 3 ex. CKAO. — SE Sulawesi, Is- land Buton-Wakarumba, 3.-7.11.1994, leg. I. JEnı$ & M. Srrsa, 3 ex. ZSM, 1 ex. SMNS. - SE Sulawesi, Airport 30km W Kendari, 11.-14.11.1994, leg. I. JEnı$ & M. Srrpa, 1 ex. ZSM. - S Sulawesi, Puncak Palopo, V.1999, local collector, 8 ex. CSBC. - Central Sulawesi, 15-25 km S Pendolo, Mayoa, 7.-10.1V.1999, leg. S. BECVAR & P. ZABRANSKY, 2 ex. CSBC. - Central Su- lawesi, 35 km NW Palopo, 1400 m, 18.IV.1999, leg. S. BECvAR & P. ZABRANSKY, 5 ex. CSBC. — Bali, Danau Buyan, 1300 m, 19.-21.11.1994, leg. L. Borm, 2 ex. SMNS. - W Bali, Mt. Bahukaru, 1100m, 26.X.2005, leg. E. Hess, 3 ex. CRGT. - Flores, Moni, 800m, 31.1.-1.11.2002, leg. M. HOFFMANN & R. RicHter, 1 ex. CRGT. Etymology: Named after the island Sumbawa, where parts of the type series were col- lected. Description: Dorsal view see Fig.72, dorsal side dark ferrugineous without metallic shine, body length 6.0-7.8mm. Genae not broader than eyes, frons without distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of pronotum slightly protruding, lateral margin distinctly sinuated, pronotal disc with rough punctation forming confluent longitudinal wrinkles and with medial impres- sion, between punctures with granules. Alternate elytral intervals 3, 5, 7 with higher and confluent longitudinal granules forming keels, intervals 2, 4, 6 with somewhat lower and more separate longitudinal granules. Tibiae in both sexes externally with- out distinct keels. Aedeagus see Fig. 135. Diagnosis: Bradymerus sumbawaicus n.sp. shares with B. elongatus (Perty, 1831) from Java the undilated genae and the general structure of the elytra with the higher alternate intervals, but can be separated by a different shape of the pronotum (Figs.27, 72; broader in B. elongatus), by a different structure of the pronotum (Figs. 27, 72; more convex and punctation confluent with longitudinal wrinkles in 2. sumbawaicus n. sp., flat and punctation confluent but without wrinkles in B. elonga- tus), and by a different shape of the aedeagus (Figs. 92, 135). See also B. kodadai n. sp. from Borneo. 58 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Bradymerus thailandicus n.sp. (Figs. 71, 136) Holotype (4): NE Thailand, Loei Prov., Phu kradung NP, 1000 m, 6.1.2000, leg. L. BROSKIEWICZ, SMNS. Paratypes: Same data as holotype, 7 ex. SMNS, 2 ex, HNHM.-N Thailand, Chiang Mai Prov., San Pakia, 1400 m, 1.-15.V.1998, leg. V. KuBAN, 8 ex. NHMB, 3 ex. SMNS. - N Thai- land, 56km NW Chiang Mai, 7.-14.V1.1995, leg. M. SNIZEK, 2 ex. CSBC. - NW Thailand, 80 km S Mae Hong Song, Khun Yuam, 27.-31.V.1996, leg. F. KANTNER, 1 ex. CSBC. — Thai- land, Lansang NP, 500m, 18.-24.IV.1991, leg. D. KrAr, 1 ex. NHMB. —- NW Thailand, N Khun Yuam, Ban Nong Pha Koa, 20.X1.1998, leg. R. Grimm, 1 ex. CRGT. - NW Thailand, Mae Hong Song Prov., Ban Huai Po, 1600-2000 m, 9.-16.V.1991, leg. J. HorAx, 1 ex. ZSM. - Laos, Giranville, 3.-24.X1.1919, leg. R. V. DE Satvaza, 5 ex. BMNH (denticeps det. GEBIEN). — Vietnam (labelled as Annam), Da Lat (labelled as Dalat), 29.1II.1924, leg. R. V. DE Satvaza, 1 ex. BMNH. -N Vietnam, Quang ninh Prov., Halong, 29.V.-1.V1.1985, leg. J. Picka, 12 ex. MHNL, 2 ex. SMNS. — N Vietnam, Hai Phong, Halong (labelled as Han-Long), 31.V.1985, lex. GSBC, Etymology: Named after Thailand, where most of the types were collected. Description: Dorsal view see Fig.71, dorsal side dark ferrugineous without metallic shine, body length 5.8-7.8 mm. Genae broader than eyes, frons with distinct supraorbital keels. Last 6 antennomeres forming a club. Anterior corners of prono- tum protruding, lateral margin with distinct crenulation, pronotal disc with rough punctation and without medial impression, between punctures with granules. Ely- tral interval 1 (only near the scutellum) and alternate intervals 3, 5, 7 with distinct keels, keels with small granules. Tibiae in both sexes externally with distinct keels; in males posterior tibia internally near the tip with a row of distinct long, yellow setae. Aedeagus see Fig. 136. Diagnosis: In external morphology, Bradymerus thailandicus n. sp. is very sim- ilar to B. crenulicollis and B. serricollis. These three taxa share the broad genae, the rough pronotum with protruding anterior corners and crenulated lateral margin and the elytra with distinct alternate keels. However, B. thailandicus n. sp. can be recog- nized by the comperatively long and parallel joint elytra, by the nearly naked elytral intervals (with several broader setae in B. crenulicollis and B. serricollis), and mainly by the completely different shape of the aedeagus (Figs. 91, 132, 136). 5 Doubtful taxa of Bradymerus The following taxa must remain as nomina dubia, because the descriptions are too poor and the corresponding types are not available for examination. It might be even possible that these taxa do not belong to Bradymerus but to different genera, for ex- ample to Gonocephalum Solier, 1834. Bradymerus elegans Pic, 1926 Indochina Bradymerus indosinensis Gebien, 1939 Indochina Bradymerus seminitidus Pic, 1926 (homonym of B. seminitidus Arrow, 1900) Bradymerus simplicithorax Pic, 1926 Indochina Bradymerus spinicollis Fairmaire, 1896 India Bradymerus viridescens Pic, 1926 Indochina Bradymerus vitalısi Pic, 1926 Cambodia SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 59 6 Transfer to the genus Derosphaerus Derosphaerus opacicollis (Kulzer, 1951) n. comb. Bradymerus opacicollis Kulzer, 1951. Studied type material: Philippines, Island Sibuyan, leg. C. F Baker, ¢ holotype NHMB-E Remarks: When describing Bradymerus opacicollis, KULZER (1951) pointed out, that several characters do not fit whithin Bradymerus, for example the long anten- nae, the long shape of the tarsomeres and the shapes of the pronotum and proster- num; and that this taxon therefore very probably represents a new genus. These are all characters of the species-rich genus Derosphaerus Thomson, 1858 (syn. En- cyalesthus Motschulsky, 1860), as well as the longer ventral setation of the tar- someres, thus, this taxon is herewith transferred to this genus. With 8.5 mm body length, however, it represents a relatively small species among the congeners. The holotype is a female and not a male as stated by KuLzerR (1951). Distribution: Philippines (Island Sibuyan). 7 Identification key for the Oriental species of Bradymerus This key is compiled only as a help for identification and not for phylogenetic interpreta- tions, it does not include all diagnostic characters and is suitable only for males because of the use of male sexual characters. In a few species only females are known as yet and the existence or lacking of an armed anterior tibia in males is tentatively concluded from related species. Not included in the key is Bradymerus grandis Fairmaire, 1893 because of its doubtful status. 1 Dorsal side with distinct or at least (in 2 species) with a feeble metallic shine ......... 2 =) sJorsalsideswathoutmetallic-s hiner re en men eee ea ae ede es 13 2 Larger species, body length 8.5-12.0 mm. Male anterior tibia without armature ....... 3 — Smaller species, body length 7.5-8.3 mm. Male anterior tibia at the ventral side with tooth CASAS 10,3 les Loe eg oth iy ut EN er eRe tn MR EM BT Res HN ER 12 3° ‘Last’ 4 antennomeres forming a club (as in Fig.31) 2... mn ee eee bee ete 4 — Last 5 antennomeres forming.a club (as: in Fig:32) 2.02842. 5 540 de ee et 22. ln bes 5 4 Anterior corners of pronotum protruding; pronotal disc with confluent punctation; alter- nateselytralinfervals.-with.keels tFıg-17) ra. an ee B. crassicollis — Anterior corners of pronotum not protruding; pronotal disc with fine and separate punc- tation; elytral intervals without any keels (Fig.15)................... B. caeruleipennis 5 Dorsal side only with a feeble metallic shine (Fig. 48). Aedeagus as in Fig. 119. — Endemic species of the small island Lan Hsu near Taiwan ................. B. masumotoi n. sp. — Dorsal side with distinct metallic shine. - Different distribution ................... 6 6 Elytral intervals convex without any keels or'granules .....6.4.0.00080 00s 002... 7 — Elytral intervals at least partly with keels or granules ....... 00... cece eee eee eee 9 7 Anterior corners of pronotum distinctly protruding (Fig.47) .......... B. laevicostatus — Anterior corners of pronotum not protruding ......... 0... c cece eee eee ee teens 8 8 Pronotum convex with rounded lateral margins (Fig. 24). - Sri Lanka... B. cyaneipennis — Pronotum flat with parallel lateral margins (Fig. 43). Aedeagus as in Fig. 106. — Philippines sn Dest Th ier OMe en Be ah BS Eee ORs Sas ar RES wall EO cl BE lic B. tris 9 Internal elytral intervals with smaller, external intervals with bigger separated granules I ee RE, ele MO NR ae eben L ieee Ment. l a, mates, fs ee ae ena B. violaceus — Elytral intervals at least partly with keels, without granules ..................0004 10 10 Only external elytral intervals with feeble keels (Fig. 19). Aedeagus as in Fig.85 ........ eee OL a SERRE ee ee Se DE re OL ee Pe ee Oe er See B. carinatus 60 11 12 13 14 15 16 17 18 19 20 21 22 23 24 25 26 27. STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Lateral margin of pronotum distinctly sinuated before posterior angles (Fig. 5). - Sri Lan- Fea Pe pire io es eee Sage Sadia Mer ee bd ag nee Sate ean ote a u ienle B. aeratus Lateral margin of pronotum not sinuated before posterior angles (Fig. 28). Aedeagus as in Rigs 95s IR bi lippimestiac a a ne oat seine Ese Me B. eschscholtzi Last 3 antennomeres forming a club; elytral intervals slightly convex; anterior corners of pronotum-not protruding. (Eid) ng ac are ne Reet eee B. aeneus Last 4 antennomeres forming a club; elytral intervals flat; anterior corners of pronotum protruding (Fig. 63). Aedeagus as in Fig.127 .................00 08. B. reibnitzi n. sp. Anterior tibia in males with 1-2 distinct teeth at the inner side (as in Fig.31) ........ 14 Anterior tibia in males without any armature’... 2. eek cee cae ser ne 24 Anterior tibia in males with a pair of distinct teeth at the inner side (Fig. 31). Aedeagus as I CAG) eer. ea i ee ODE Each Ree EIERN EN ee te B. fouquei n. sp. Anterior tibia in males with a single tooth at the inner side ..........---000 sees eee 15 Loothanythesnidd erat the calle tibia, pve. Sa u a eh eee Oa ele 16 Tooth in the antenor'third of the male tibia®. „2... u... san see ea 17 Body length 9.0mm; pronotum longer; alternate elytral intervals 3, 5, 7 without distinct keels and with feeble pore-bearing granules (Fig. 22). Aedeagus as in Fig.88 ........... Bde Be eet Sa Sa ue et es ee reer ee B. crockerensis n.sp. Body length 6.5-7.2 mm; pronotum wider; all elytral intervals with a row of distinct pore- bearing granules (Fig. 35). Aedeagus as in Fig.101 ..............0000- B. grimmi n. sp. Last 3 antennomeres forming a club; antennae extraordinary long (Fig. 45). Aedeagus as in Hig MUO fund tere ee N LE Nal eed Eh 1 wae B. kinabalicus n. sp. Last 4 antennomeres forming a club; antennae of normal length................... 18 Dorsal’side- dull blackish 22 Je: 2.4, 00. 20. 0 ne nen de Be de pees ae ne 19 Dorsal side shining blackish, brownish or ferrugineous ............. 00. c seer ee eee 20 Anterior corners of pronotum protruding; only alternate elytral intervals 5, 7 with keels; punctures of the elytral rows deeply impressed, longitudinal (Fig.58). Aedeagus as in POT a N NEE hd TE ET ee B. merkli n. sp. Anterior corners of pronotum not protruding; alternate elytral intervals 3, 5, 7 with keels; punctures of the elytral rows normally impressed, round (Fig.49). Aedeagus as in PUG AGL means ae sen bale Peat tad ald tnd ok end, ne ER aye ne B. laoticus n. sp. All elytral intervals slightly convex and without any keels or granules (Fig. 10). Aedeagus ASAIO! FUSER Sc” A Be Be a ae eae etm Om He EU Se Se EN a B. atronitens At least some elytralintervals.with:keels:or cranules\ncar 4% ut srl lern 2 ne 21 Pronotal disc with rough and confluent punctation, between punctures with distinct granules (Fig.29). Aedeagus as in Fig.96 =....2.2.. es cat een ernennen B. fukiensis Pronotal disc with dense or even confluent punctation, but without granules........ 22 Alternate elytral intervals 3 (interrupted in the middle), 5, 7 with keels; keels with distinct granules, intervals 1, 2, 4, 6 with a row of distinct granules (Fig.36). Aedeagus as in PIG M02 Baile, ong ow ole mc nut en tnd bg eed ol ORE ON eher ee Tes B. hauseri n.sp. Elytral:stru@mure-dikterent te. RE N Se a a TE BR 23 All elytral intervals convex, interval 7 somewhat higher and with a row of fine pore-bear- ing granules (Fig. 67). Aedeagus as in Fig. 130 2. nc. u ss see nn B. sijthoffi Alternate elytral intervals 3, 5 with feeble keels, interval 7 with a more distinct keel, inter- vals 2, 4, 6 slightly convex and without any granules (Fig. 26). Aedeagus as in Fig. 89 EIER. Mena ah a rn AR BO BRET, cen 2 B. drescheri Genae narrower than ‘eyes’ 2. ya untere 8 gy 9 Ale te end ae. 4's Pee PEER Ee en 41 Genae distinctly broader than eyes or at least as broad as eyes ......... eee eee ee 25 Allelyttal intervals, with-identical fine keels2 u... rn. 22 8 2 ann AE 26 Only alternate elytral intervals with keels or intervals partly with granules instead of keels, or without keels but with rows of granules ....... 2.2.0... ne een 27 Pronotal disc with confluent punctation, between punctures without granules; tibiae ex- ternally with fine keels. Habitus as in Fig. 68. Aedeagus as in Fig. 131 ........ B. spretus Pronotal disc with dense punctation, between punctures with fine granules; tibiae exter- nally without keels. Habitus as in Fig. 61. Aedeagus as in Fig. 125 ....... B. propinquus Alternate elytral intervals 3, 5, 7 with distinct, complete and identical keels, other intervals withror without-tows'obgranuless N yon ee ALCOR, WR ees N a We es 28 Alternate elytral intervals different, mostly with granules, granules partly confluent . . 36 28 29 30 31 32 33 34 35 36 37 38 39 40 41 42 43 44 45 SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 61 Pronotal disc with a pair of prominent tubercles (Fig.57). Aedeagus as in Fig. 122 ...... ee ee re EOE ae an en ES ee eee B. nodicollis Pronotal disc without a pair of prominent tubercles 2... u... eee cece es 29 Pronotal lateral margin without distinct crenulation (as in Fig. 19) ................. 30 Pronotal lateral margin with distinct crenulation (as in Fig. 18) ..........22222220.. 31 Tibiae externally with feeble keels. Habitus as in Fig. 30. Parameres of aedeagus spade-like CEIGPOS Vesey et then cherie eee ee hes Vee BER a i, ER em B. ferruginipes Tibiae externally without keels. Habitus as in Fig.40. Parameres of aedeagus more acute EIS LOA): Sa Wh dar Pee ROS. Beech Ate Re RR MA EAN SR B. interstitialis Pronotal disc without granules between punctures (Fig.56). Aedeagus as in Fig.116 .... pith eps A AR a Eee twas te al TR NE ad ern Ot tae B. mindanaensis Pronotal disc with granules between 'punctüres.. „u 2. nennen nah nen 32 Frons without suprserbitalkeelsen en a. tet ie en ee hte ots 33 Frons with distinct supraorbitäl.keels „a aa era psec ee 34 Body length 5.3-6.0 mm (Fig. 21). Parameres of aedeagus with rounder tip (Fig.90) ..... PERS Ne Ba a En ht UW pe NT er A are WR art GAR MAR, Sento on Ne Ae Ie B. difficilis Body length 7.5-8.0 mm (Fig. 20). Parameres of aedeagus more acute (Fig. 86) ......... beet tile ake, a eh oer mie Ae EN OO PAE Canto a 2 ek ae tg B. celebensis Elytra long and parallel; elytral intervals without setation (Fig.71). Joint parameres of aedeagus pentasonal (HAs TG sie Bet tc Meee ty ee eae ae ant at tg B. thailandicus n. sp. Elytra shorter; elytral intervals with setation. Joint parameres of aedeagus triangular .. . 35 Elytral intervals with broader scale-like setae (Fig. 66). Joint parameres of aedeagus trian- sular-withroundertip (Lig 132m ce. nn ee ae We tt Mice a Mie ne wets B. serricollis Elytral intervals with thinner scale-like setae (Fig. 18). Joint parameres of the aedeagus withisinuated and acute op lig: I wits 2252 52% oh tk iene ee ee B. crenulicollis Tibia externally without distinct keels. Habitus as in Fig. 39. Aedeagus as in Fig. 108 .... Ee ee ee ee es ree i ee B. kabakovi ‘Tibravexternally swath distinct Ieee geo... Alte ee ont nage ete 37 Bronsewit aidistinct supraocbital keelsu inte ann na ae eyes omen en, 38 Kronsswithoutisuptraorbitalkkeels Han a a a Dr a Bees 39 All elytral intervals with longitudinal and confluent tubercles forming interrupted keels, alternate intervals 3, 5, 7 slightly more prominent (Fig. 32). Aedeagus as in Fig.98 ...... U IE Re Br We a a ee a Nee eel Rd EEE B. gebieni n. sp. Internal elytral intervals smooth with small granules, external intervals with prominent longitudinal granules (Fig. 9). Aedeagus as in Fig.79 ..........-0e esses eens B. aratus Body length 3.5 mm; pronotal disc between punctures with distinct tubercles; elytral in- terval 3 in the middle with prominent keel (Fig. 69). Aedeagus as in Fig. 134 ........... EN ER ems tem ee eve ci aed TE Here nen onstage ee use RM hank ee Sed B. sprecherae n. sp. Body length 4.5-7.0 mm; pronotal disc between punctures with granules; elytral structure differentd.. en... nennen Oe Oe Dalene ang me. wees mich a here Ee her Melt pes 40 Internal elytral intervals with separated granules, external intervals with longitudinal, partly confluent granules (Fig. 34). Joint parameres of aedeagus broader (Fig.100) ...... ed tN ea ae es En En et en res, ae B. granulipennis Elytral intervals with distinct longitudinal granules, on intervals 3, 5 partly confluent and forming keels (Fig. 11). Joint parameres of aedeagus narrower (Fig. 80) ........ B. asper All eytral intervals slightly convex, neither with keels nor granules ................ 42 At least some elytral intervals with distinct keels or rows of granules .............. 47 Anterior corners of pronotum protruding (as in Fig 12) ........ eee eee eee eee 43 Anterior-corners of pronotuin not protruding? u... mn. an ann NBR a 44 Body length 10.8 mm; pronotum widest in the distal third (Fig. 12). Aedeagus as in Fig. 82 ond ab ace Me ay onthe gE aaa hs. ranted ath se, Sheen pe Reon B. aulacopterus Body length 6.5-7.5 mm; pronotum widest at the base (Fig. 38). Aedeagus as in Fig.105 . MI ES oct Mee esa eat eg wee GoM AE OM eth Migs RE cae tA eat RN B. incostatus TastHantennomeresstormme.#club®3 un sate tas tos te ra oe OE ce aeons 45 Last-5antennomeres forming a. cluübr. ur. fal eens tte ee oh ee eae 46 Habitus as in Fig. 7. Joint parameres of aedeagus broad with blunt tip (Fig.77) ......... Ce ee Tee nie NER nd re ka ea ie eee ee B. andamanus Habitus as in Fig. 62. Joint parameres of aedeagus long with rounded tip (Fig. 126) ..... ee en RL Meter gez Meese SEHR ler ae Ne ul B. pseudomalayicus n.sp. 62 46 47 48 49 50 51 52 53 54 55 56 57 58 59 60 61 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Pronotum widest in the middle; pronotal disc with fine and separate punctation (Fig. 51). PCGCACUSAS LoS 44 rn denen a Te ry Eterm ates B. malayicus n. sp. Pronotum widest before middle; pronotal disc with dense and confluent punctation (Fig. 41); Aedeagus-asan Fig-104 7 ..0 2 „u Inne B. kanchenjungicus n.sp. Anterior margin of pronotum medially with striking prolongation covering completely the head; clypeus bent upwards (Fig.23) . 2.2... cc ees 2er eee weee’ B. cucullatus Anterior margin of pronotum not distinctly prolonged; clypeus not bent upwards ... 48 Lateral margin of pronotum widely separated and distinctly bent upwards (Fig. 64). PAGO US aS IIe ICID Gln, Bun a eal May nk ars ania Maite Pete eae BER BR B. riedeli n. sp. Lateral margin of pronotum not distinctly separated and not bent upwards ......... 49 Frons with deep and extraordinary wide supraorbital furrow; pronotum humped (Biss). Acdeasusasın FAZIT net cates te 1c he bee B. michihikoi n. sp. Frons at most with deep, but never with wide supraorbital furrow; pronotum at most convex-but:not’hüumped. (ey nto 28. Se eae ee a u inet eee 50 Anterior corners of pronotum not protruding ......... 0... cece cece eee eee eee 51 Anterior corners of pronotum distinctly or at least slightly protruding (as in Fig. 12) ...54 All elytral intervals wrinkled by rough punctation, internal intervals flat and anteriorly with a row of distinct granules, external intervals with indistinct keels (Fig. 46). Aedeagus ASTI US LAD Pr. ns BE Mean UR ae eels te gee Mee ete estate cs na ase peg asc B. kulzeri n.sp. Structure:of\the elytral interyals.ditferent nr ee ee ee ar 52 Alternate elytral intervals 3, 5, 7 with keels, intervals 2, 4, 6 smooth and without punctures and granules (Fig:6)..Aedeagus as in Fig.76. 2... un en Lena deat sen B. alternicostis Structure.of the elytral intervals different 4... cic. nun. nn ee eee 53 Pronotum widest near the anterior margin; elytral intervals 3 posteriorly, 4 posteriorly, 5, 6, 7 completely with keels; internal intervals 1-4 flat and without granules (Fig. 52). Aedeaous-as im: Pisa TPB on tue Pao Pet ie aide dete ra A ced, B. majeri n. sp. Pronotum widest in the middle; alternate intervals 3 anteriorly, 5 and 7 completely with keels; internal intervals 2, 4, 6 with separated distinct granules (Fig. 59). Aedeagus as in Pig: TD ARIAS ang en Eee ae tit, Se Ul rc hansyanch ars um tena B. pertyi Elytral intervals mostly with isolated granules (as in Fig.53) 1.1.0... 00. eee eee eee 55 Elytral intervals mostly with keels or with longitudinal confluent tubercles forming keels be ees oe res at Dy LN alg LO BE I Sows EURE BER TER FRE Rs eh 28 We baud Pi bwin en. otis eae 58 Pronotumawaidest.in the-anterior thirds, 20.2.2... 3 he a Sees oa 56 Pronotuin widest nt thermiddle'tr. Wirt atch ce Moet BAS ORLA Por De LR TL 57 Body length 7.8-9.0 mm; internal elytral intervals 1-3 with a row of separate granules, ex- ternal intervals with confluent longitudinal granules (Fig. 8). Joint paramers of aedeagus trianeular with.acute tips Fig «7S u. EIN + x's atin tive. nen lee B. andoi n. sp. Body length 7.2 mm; elytral intervals with distinct granules, granules on intervals 5, 7 sometimes confluent (Fig.53). Joint parameres of aedeagus finger-like with rounded tip CBig ALS). ansehen en ek eat ee tad te eae ended B. maramagicus n. sp. Last 5 antennomeres froming a club; anterior corner of pronotum extremely protruding, acute (Fig. 3). Joint parameres of aedeagus finger-like with rounded tip (Fig. 74) ........ en ER 1 ne na ee ea ee Be er a, ERO ae gd er B. acutangulus Last 6 antennomeres forming a club; anterior corner of pronotum not extremely protrud- ing, round (Fig. 33). Joint parameres of aedeagus triangular with rounded tip (Fig. 99) ... Poe ne a a EEE Sal og Ae eM, pace AR OG 2 Pen as en ave B. gerstmeieri n. sp. All elytral intervals with identical keels or only internal two intervals with traces of keels but-alltotherintervalsavithidentical Keelsae tm EA ces eee Bil ies be Fee 59 Keels only present on alternate intervals or in other pattern, but not identical on all inter- Vals nee AE Sedat a LE Rk oR AY RO RENDER a condi na a et Be ARE ae, ee 62 AlPelytral intervals:wath- identical: Keel sav. 2. Aue nee nn a tite van 60 Only internal intervals 1, 2 with traces of keels but all other intervals with identical keels heh dente NED DE ORT FR N) > CNS I gt geaNGE LS. Sent ered SOMOS SOY Fee RONG AR Vt gt hie tee 61 Body length 7.0-6.5mm; pronotum longer (Fig.37). Joint parameres of aedeagus with acute.bip (ie 1) RS ea .. 1 Me BE nn RE Be aint en otic a B. impressicollis Body length 5.3-6.5 mm; pronotum wider (Fig.2). Joint parameres of aedeagus with koundedstipr lien 75 en re ne not Br Ne Ta gk ap, CM LD a ae B. acuticostis Body length 7.5 mm; joint elytra long (Fig. 65). Basal piece of aedeagus long (Fig. 129). — Endemic species of Christmas Island . 2... ..s 00000. eee eet evens B. seminitidus SCHAWALLER, ORIENTAL SPECIES OF BRADYMERUS 63 — Body length 5.0-7.0mm; joint elytra shorter (Fig.16). Basal piece of aedeagus short (Ei Widespread’ species. sw inastesuee arora bron he et nis ne ame eons B. clathratus 62: Last S-afitenmomieres tormiune a Club. es con! Taal peter td tee Cae es 63 = Last -6-antennomeres toring arcluber so HAN EN eee AM en stat sts 66 63 Body length 6.0 mm; anterior corners of pronotum extremely protruding, acute (Fig. 70). Pedeaous a8 in) Fig 133. are hate ee A ee ee epee the eae hime B. sumatranus n. sp. — Body length 6.5-9.0 mm; anterior corners of pronotum not extremely protruding, round BEIN. Bath Sa oR HANSE N RIES tina) GE RE En bee Re LS tm de A eg IAN GAS hus Coe sup a m Bl BEA ek 64 64 Pronotum widest in the middle; alternate elytral intervals 3 (only anteriorly), 5, 7 with keels, intervals 2, 5, 7 flat and without granules (Fig. 13). Aedeagus as in Fig.81 ........ pss ee aye etm ae Bk ord ele cee Sat ee oh Rae ooo AEM Rc A a st B. bifurcatus — Pronotum widest in the anterior third; structure of elytral intervals different ........ 65 65 Alternate elytral interval 3 posteriorly with a low keel, intervals 5 and 7 with complete and high keels, keels with granules (Fig. 14). Aedeagus as in Fig. 84 ..... B. bocakorum n. sp. — Elytral intervals 3 posteriorly, 5 and 6 anteriorly, 7 completely with keels, keels without pramules(hie260): ra see M nici e ct Mee ae PRA Se A la ler Bea as B. planicollis 66 Alternate elytral intervals 2, 4, 6 only with a few and feeble granules (Fig. 54). Aedeagus as TAGs 1200, ee tet ne a a ants aie es een B. mcgregori — Alternate elytral intervals 2, 4, 6 either with keels or granules in a different pattern ... 67 67 Small species (body length 5.5-7.0 mm) with long and parallel elytra (Fig. 25). Aedeagus as I EIS IE oe eed ROE eres fo, Perens Peters DERE PE BE, B. emasicus n. sp. — Somewhat larger species in the average (body length 6.0-9.0 mm) with broader and more compact elytra ....... B. elongatus (Java), B. kaszabi n.sp. (Sulawesi), B. kodadai n. sp. (Borneo), B. lombokicus n.sp. (Lombok), B. sumbawaicus n.sp. (Sumbawa, Sulawesi) [The identification of these five quite similar species (mostly endemic insular populations) is only possible by a fastidious comparison of shape and structure of the pronotum, pat- tern of the keels and granules on the elytral intervals, and shape of the aedeagus; see the details in the descriptions and diagnoses. 8 References BOUCHARD, P., LAWRENGE, J. F., Davies, A. & NEwTon, A. F. (2005): Synoptic classification of the World Tenebrionidae (Insecta: Coleoptera) with a review of family-group names. — Annales Zoologici 55: 499-530. FAIRMAIRE, L. (1893): Coléoptéres nouveaux des Indes orientales, de la famille des Scarabaei- dae, Rhipidoceridae, Tenebrionidae et Oedemeridae. — Notes from the Leyden Muse- um 15: 17-64. GEBIEN, H. (1913): Die Tenebrioniden der Philippinen. — Philippine Journal of Science 8: 373433. GEBIEN, H. (1925): Die Tenebrioniden (Coleoptera) des indomalayischen Gebietes, unter Berücksichtigung der benachbarten Faunen, III. Die Gattungen Bradymerus, Chaetop- sia, Danodema, und Dicraeosis. - Philippine Journal of Science 26: 535-576. GEBIEN, H. (1936-1944): Katalog der Tenebrioniden (Col. Heteromera). — Part 1 (1936): Pub- blicazioni del Museo entomologico Pierro Rossi 2; Part 2 (1938-1942): Mitteilungen der Münchener entomologischen Gesellschaft 28-32; Part 3 (1942-1944): Mitteilungen der Münchener entomologischen Gesellschaft 32-34. Kaszas, Z. (1954): Uber die von Herrn J. KLApPERICH in der chinesischen Provinz Fukien gesammelten Tenebrioniden (Coleoptera). — Annales historico-naturales Musei natio- nalis hungarici 5: 247-264. Kaszas, Z. (1980): Angaben zur Kenntnis der Tenebrioniden Nordvietnams (Coleoptera). — Annales historico-naturales Musei nationalis hungarici 72: 169-221. Kuwzer, H. (1951): Fünfter Beitrag zur Kenntnis der Tenebrioniden. — Entomologische Ar- beiten aus dem Museum G. Frey 2: 461-573. KULzer, H. (1957): Insects of Micronesia. Coleoptera: Tenebrionidae. - Insects of Micronesia 17: 185-256. SHIBATA, T. (1980): Notes on Tenebrionidae from Taiwan and Japan, III. (Coleoptera). - Entomological Review of Japan 34: 63-74. 64 STUTTGARTER BEITRÄGE ZUR NATURKUNDE Ser. A, Nr. 694 Author’s address: Dr. WOLFGANG SCHAWALLER, Staatliches Museum für Naturkunde, Rosenstein 1, 70191 Stuttgart, Germany; e-mail: schawaller.smns@naturkundemuseum-bw.de Manuscript received: 21.X1.2005, accepted: 9.1.2006. ISSN 0341-0145 Autoren-Richtlinien: http://www.naturkundemuseum-bw.de/stuttgart/schriften Schriftleitung: Dr. Hans-Peter Tschorsnig, Rosenstein 1, 70191 Stuttgart Gesamtherstellung: Gulde-Druck, 72072 Tübingen